張艾英,郭二虎,刁現(xiàn)民,范惠萍,李瑜輝,王麗霞,郭紅亮,程麗萍,吳引生
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2005—2015年西北春谷中晚熟區(qū)谷子育成品種評(píng)價(jià)
張艾英1,郭二虎1,刁現(xiàn)民2,范惠萍1,李瑜輝1,王麗霞1,郭紅亮1,程麗萍1,吳引生1
(1山西省農(nóng)業(yè)科學(xué)院谷子研究所/特色雜糧種質(zhì)資源發(fā)掘與育種山西省重點(diǎn)實(shí)驗(yàn)室,山西長(zhǎng)治 046011;2中國(guó)農(nóng)業(yè)科學(xué)院作物科學(xué)研究所,北京100081)
對(duì)11年來(lái)西北春谷中晚熟區(qū)育成的谷子品種產(chǎn)量、農(nóng)藝性狀、抗逆性及品質(zhì)性狀綜合分析,為西北春谷中晚熟區(qū)谷子新品種選育、推廣及資源利用提供理論參考。基于2005—2015年國(guó)家谷子品種區(qū)域試驗(yàn)西北春谷中晚熟區(qū)數(shù)據(jù),對(duì)參試品種及通過(guò)鑒定品種的產(chǎn)量、主要農(nóng)藝性狀進(jìn)行比較與分析。并對(duì)通過(guò)鑒定的30個(gè)品種抗逆性、品質(zhì)進(jìn)行分析。2005—2015年參試品種及通過(guò)鑒定品種的產(chǎn)量有逐年增加的趨勢(shì)。參試品種和通過(guò)鑒定品種的農(nóng)藝性狀在年度間存在一定的變異,參試品種的株高、單穗重、穗粒重、出谷率有逐年增加的趨勢(shì),生育期有縮短的趨勢(shì);通過(guò)鑒定的30個(gè)品種和對(duì)照相比,生育期、株高有降低趨勢(shì),株高變幅為105—165 cm;穗長(zhǎng)在17—27 cm,70%的品種分布在19—23 cm;單穗重變幅為15—25 g,80%品種為18—22 g;穗粒重為12—20 g;出谷率分布在74%—84%,80%的品種在75%—80%;千粒重變異范圍較大,分布在2.5—3.4 g,有9個(gè)品種的千粒重超過(guò)對(duì)照,公頃穗數(shù)分布在33萬(wàn)—43萬(wàn)。對(duì)通過(guò)鑒定品種農(nóng)藝性狀與產(chǎn)量的相關(guān)分析表明,產(chǎn)量與單穗重、穗粒重和公頃穗數(shù)呈顯著正相關(guān),與生育期呈負(fù)相關(guān)。鑒定品種總體抗性有提高的趨勢(shì),其中抗倒性明顯優(yōu)于對(duì)照,紅葉病和白發(fā)病為主要病害,谷銹病、谷瘟病、紋枯病、蛀莖率也有不同程度地發(fā)生。穗松緊度多為中等偏緊類型,熟相中等偏好為主。品質(zhì)上,通過(guò)鑒定品種的米色全為黃色,優(yōu)質(zhì)米的粗蛋白、粗脂肪含量及膠稠度偏低,賴氨酸含量相對(duì)較高。鑒定品種中包括了糯質(zhì)、高蛋白、高脂肪及糧草兼用、抗除草劑、優(yōu)質(zhì)米等多種類型,豐富了品種類型。西北春谷晚熟區(qū)谷子品種的選育取得了一定的進(jìn)步,所育品種在數(shù)量、多樣性、產(chǎn)量、品質(zhì)等方面有了一定的提高,但沒(méi)有取得大的突破。在育種方法上比較單一,多采取簡(jiǎn)單雜交、系選的方法,應(yīng)根據(jù)育種目標(biāo)要求,開(kāi)展材料創(chuàng)新,融合回交、復(fù)交、理化誘變等多種方法,并結(jié)合分子育種新技術(shù),提高育種水平。在育種目標(biāo)上,重點(diǎn)培育品質(zhì)優(yōu)良、矮稈抗倒、生育期略短、適合機(jī)械化收獲的品種,培育多種類型的抗除草劑品種,滿足谷子規(guī)?;a(chǎn)的需求。
春谷;中晚熟區(qū);區(qū)域試驗(yàn);農(nóng)藝性狀;相關(guān)分析
【研究意義】谷子((L.) Beauv.)主要分布于中國(guó)東北、華北和西北地區(qū)[1-2],由于生態(tài)環(huán)境的差異形成了適合不同生態(tài)區(qū)種植的品種。西北春谷中晚熟區(qū)是谷子生產(chǎn)的主要產(chǎn)區(qū),也是重要的優(yōu)質(zhì)米生產(chǎn)區(qū)域,著名的貢米“沁州黃”產(chǎn)于本區(qū)的沁縣。該區(qū)小米中蛋白質(zhì)、脂肪含量中等,但適口性好,對(duì)短日照反應(yīng)中等至敏感,對(duì)長(zhǎng)日照反應(yīng)中等,溫反應(yīng)不敏感[3-5]。通過(guò)對(duì)多年來(lái)西北春谷區(qū)區(qū)域試驗(yàn)數(shù)據(jù)的匯總分析,可以客觀、全面地了解本區(qū)域谷子品種變化情況及整體育種水平,研究分析影響谷子產(chǎn)量主要構(gòu)成因素,為本生態(tài)區(qū)新品種評(píng)價(jià)、確定未來(lái)育種目標(biāo)及資源利用提供理論依據(jù)[6-10]?!厩叭搜芯窟M(jìn)展】水稻[11-13]、小麥[14-16]、玉米[17]、油菜[18]根據(jù)多年區(qū)域試驗(yàn)和生產(chǎn)示范的技術(shù)資料確定階段性的實(shí)際育種目標(biāo)或育種方向。欒素榮等[19]對(duì)12個(gè)西北區(qū)谷子中晚熟品種進(jìn)行試驗(yàn)表明,影響谷子產(chǎn)量的主要因素是穗粒重、單穗重、出谷率和穗粗。王軍等[20]對(duì)2008年國(guó)家西北春谷中晚熟區(qū)區(qū)域試驗(yàn)品種的適應(yīng)性進(jìn)行了分析,選出長(zhǎng)生07、朝108、汾選8號(hào)和太選7號(hào)等適應(yīng)性和穩(wěn)產(chǎn)性較好的品種。楊慧卿等[21]對(duì)2008年國(guó)家西北春谷中晚熟區(qū)區(qū)域試驗(yàn)品種的相關(guān)和通徑分析認(rèn)為,生育期、穗粒重和單穗重是構(gòu)成西北春谷中晚熟區(qū)谷子單株產(chǎn)量的主要因素,其次是株高和出苗—抽穗天數(shù)。秦嶺等[22]對(duì)不同生態(tài)區(qū)60份谷子品種的12個(gè)主要農(nóng)藝性狀與產(chǎn)量相關(guān)性分析得出單株穗重、主穗直徑對(duì)產(chǎn)量的直接效應(yīng)較大。劉敏軒等[23]對(duì)中國(guó)谷子育成品種維生素E含量分布規(guī)律及其與主要農(nóng)藝性狀和類胡蘿卜素的相關(guān)性分析得出,中國(guó)谷子VE總量與類胡蘿卜素含量呈顯著正相關(guān),與千粒重呈顯著負(fù)相關(guān)。孫宇燕等[24]對(duì)春谷農(nóng)藝性狀和品質(zhì)性狀進(jìn)行了變異分析,為春谷種質(zhì)資源和雜交親本選配提供了參考依據(jù)。以上研究在一定程度上為西北春谷中晚熟區(qū)谷子育種和資源利用提供了一定的理論依據(jù)?!颈狙芯壳腥朦c(diǎn)】前人研究匯總數(shù)據(jù)單薄、品種少,系統(tǒng)性不強(qiáng),所以結(jié)論有一定的局限性,對(duì)年度間、區(qū)域間、品種間的差異也缺少周全性。【擬解決的關(guān)鍵問(wèn)題】本研究通過(guò)對(duì)11年來(lái)西北春播中晚熟區(qū)育成的谷子品種農(nóng)藝性狀、產(chǎn)量表現(xiàn)、抗逆性及品質(zhì)性狀綜合分析,為西北春谷中晚熟區(qū)新品種推廣、選育、資源利用及谷子產(chǎn)業(yè)發(fā)展提供理論參考。
數(shù)據(jù)來(lái)源于2005—2015年西北春谷中晚熟區(qū)區(qū)域試驗(yàn)。每年承試點(diǎn)為10個(gè)左右,年度間存在一定的調(diào)整變化,其中5個(gè)試點(diǎn)連續(xù)承擔(dān)了試驗(yàn),分別是遼寧水土保持研究所、山西省農(nóng)業(yè)科學(xué)院谷子研究所、山西省農(nóng)業(yè)科學(xué)院經(jīng)濟(jì)作物研究所、山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所、陜西延安市農(nóng)業(yè)科學(xué)研究所;其他單位階段性承擔(dān)了試驗(yàn),分別是甘肅秦安縣種子管理站、北京門(mén)頭溝區(qū)清水鎮(zhèn)、河北承德市研究所、遼寧錦州市農(nóng)業(yè)科學(xué)研究所、西北農(nóng)林科技大學(xué)、遼寧風(fēng)沙保持研究所、甘肅天水市農(nóng)業(yè)科學(xué)研究所、山西沁州黃小米公司及沁縣種子站。2005—2015年有84個(gè)品種參加了127年次試驗(yàn),通過(guò)國(guó)家鑒定品種30個(gè)(電子附表1),其中一級(jí)優(yōu)質(zhì)米5個(gè),二級(jí)優(yōu)質(zhì)米11個(gè),抗除草劑品種2個(gè),糧草兼用品種一個(gè)。
1.2.1 數(shù)據(jù)匯總 對(duì)2005—2015年所有參試品種及通過(guò)鑒定品種的產(chǎn)量、主要農(nóng)藝性狀、抗逆性、品質(zhì)性狀等進(jìn)行匯總分析,并與對(duì)照承谷8號(hào)和長(zhǎng)農(nóng)35號(hào)進(jìn)行對(duì)比。
1.2.2 統(tǒng)計(jì)分析 數(shù)據(jù)分析采用DPS軟件[25]進(jìn)行,顯著性檢驗(yàn)采用新復(fù)極差法,圖表采用Microsoft Excel 2003軟件繪制。
2.1.1 參試品種產(chǎn)量變化 圖1為2005—2015年各參試品種和對(duì)照(2005—2006年對(duì)照品種為承谷8號(hào),2007—2015年對(duì)照品種為長(zhǎng)農(nóng)35號(hào))產(chǎn)量的變化情況,可以看出,參試品種公頃產(chǎn)量由2005年低于4 000 kg到2015年約5 000 kg,呈現(xiàn)逐年增加的趨勢(shì),且對(duì)照表現(xiàn)為類似趨勢(shì);參試品種產(chǎn)量各年度間存在一定的差異,2014年產(chǎn)量最高,2006—2007年產(chǎn)量最低,且最高產(chǎn)量與最低產(chǎn)量間存在顯著差異,2005—2015年參試品種產(chǎn)量均值(2010年除外)高于對(duì)照。從產(chǎn)量結(jié)果變化情況看,新品種的增產(chǎn)潛力明顯提高,2007年更換對(duì)照品種后,對(duì)提升品種的產(chǎn)量水平發(fā)揮了較好的標(biāo)桿作用。
不同字母表示P=0.05水平差異顯著。下同
2.1.2 參試品種農(nóng)藝性狀變化 2005—2015年各年度參試品種農(nóng)藝性狀及產(chǎn)量情況(表1)。由表可知,參試品種農(nóng)藝性狀年度間變異較大,株高、單穗重、穗粒重、出谷率、公頃穗數(shù)有增加的趨勢(shì),穗長(zhǎng)和千粒重年度間無(wú)顯著差異,生育期有縮短的趨勢(shì)。從主要農(nóng)藝性狀的變化情況看,春谷中晚熟區(qū)新育成品種產(chǎn)量的提高,主要靠培育大穗型品種,增加單穗重,通過(guò)縮短品種的生育期改善品種的結(jié)實(shí)性。品種株高呈先增后降的趨勢(shì),降低幅度不大,不適應(yīng)現(xiàn)代谷子的機(jī)械化生產(chǎn),應(yīng)將降低株高作為今后的育種目標(biāo)。
2.2.1 產(chǎn)量變化情況 對(duì)通過(guò)鑒定的30個(gè)品種的產(chǎn)量及產(chǎn)量標(biāo)準(zhǔn)差匯總(圖2),可以看出通過(guò)鑒定的品種產(chǎn)量有逐年增高的趨勢(shì)。通過(guò)鑒定的品種2007—2011年平均公頃產(chǎn)量4 247.01 kg,2012—2015年平均公頃產(chǎn)量為4 824.49 kg,后期平均公頃產(chǎn)量增加了577.48 kg,增幅為13.6%(對(duì)照長(zhǎng)農(nóng)35號(hào)2007—2011年平均公頃產(chǎn)量4 128.57 kg,2012—2015年平均公頃產(chǎn)量4 654.54 kg,后期平均公頃產(chǎn)量增加了525.97 kg,增幅為12.74%),總之,品種的增產(chǎn)潛力呈提高的趨勢(shì)。在所有通過(guò)鑒定的品種中,糯質(zhì)品種1個(gè)(汾選8號(hào))、滿足農(nóng)牧交錯(cuò)帶應(yīng)用的糧草兼用型品種(蒙金谷一號(hào))1個(gè)、抗除草劑品種2個(gè),一級(jí)優(yōu)質(zhì)米5個(gè),二級(jí)優(yōu)質(zhì)米11個(gè),品種類型的多樣性為滿足谷子產(chǎn)業(yè)品種多樣化的需求發(fā)揮了重要作用,同時(shí)也對(duì)品種整體產(chǎn)量水平的提高造成一定的影響。通過(guò)鑒定的品種中,增產(chǎn)幅度在8%以上的品種17個(gè),占鑒定品種總數(shù)的一半以上。
表1 2005—2015年西北春谷中晚熟區(qū)參試品種主要農(nóng)藝性狀及產(chǎn)量情況
不同字母表示=0.05水平差異顯著。下同 Different small letters within a column indicate significant difference at the 0.05 level. The same as below
圖2 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種產(chǎn)量變化情況
2.2.2 農(nóng)藝性狀變化情況 對(duì)春谷中晚熟區(qū)通過(guò)鑒定的30個(gè)品種農(nóng)藝性狀匯總。11年來(lái)參試品種生育期呈現(xiàn)逐年下降的趨勢(shì),一般在120—135 d。由于2005—2006年的對(duì)照品種為承谷8號(hào),其生育期短,所以鑒定品種的生育期都比其高,2007年對(duì)照品種換為長(zhǎng)農(nóng)35號(hào),更換對(duì)照品種后通過(guò)鑒定品種的生育期與對(duì)照相近,并呈現(xiàn)生育期縮短的趨勢(shì),近幾年的新育成品種生育期都低于130 d,在120 d上下波動(dòng)。新育成品種的株高波動(dòng)在105—165 cm。2005—2006年通過(guò)鑒定的品種株高高于對(duì)照承谷8號(hào),2007年以后只有個(gè)別品種的株高高于對(duì)照長(zhǎng)農(nóng)35號(hào),多數(shù)品種低于對(duì)照;30個(gè)品種穗長(zhǎng)分布在17—27 cm范圍內(nèi),超過(guò)23 cm的有7個(gè),低于19 cm的有2個(gè),多數(shù)品種分布在19—23 cm范圍內(nèi),2005—2006年通過(guò)鑒定的品種穗長(zhǎng)低于對(duì)照承谷8號(hào),2007年以后只有個(gè)別品種的穗長(zhǎng)低于對(duì)照長(zhǎng)農(nóng)35號(hào);對(duì)照品種和通過(guò)鑒定品種的單穗重呈逐年升高的趨勢(shì),其重量分布在15—25 g,低于18 g的有2個(gè),高于22 g的有4個(gè),大多數(shù)品種變化趨勢(shì)和對(duì)照變化一致,單穗重最低的品種為興谷66;對(duì)照和通過(guò)鑒定品種穗粒重呈逐年增加趨勢(shì),穗粒重分布在12—20 g,穗粒重最低的品種為興谷66;出谷率分布在74%—84%,低于75%的品種有2個(gè),高于80%有4個(gè),有8個(gè)通過(guò)鑒定品種的出谷率高于對(duì)照,其余的都低于對(duì)照;對(duì)照千粒重變化平緩,基本上維持在3 g左右,但通過(guò)鑒定品種的千粒重變異范圍較大,分布在2.5—3.4 g,有9個(gè)品種的千粒重超過(guò)對(duì)照,其余都低于對(duì)照,千粒重最高的品種為朝谷19,最低的品種為晉谷42號(hào);公頃穗數(shù)分布在33萬(wàn)—43萬(wàn),呈逐年增高趨勢(shì),多數(shù)品種和對(duì)照的變化趨勢(shì)一致,比對(duì)照明顯高的為燕谷16號(hào)(電子附圖1)。
2.2.3 產(chǎn)量及主要農(nóng)藝性狀的相關(guān)性分析 西北春谷中晚熟區(qū)通過(guò)鑒定品種主要農(nóng)藝性狀平均值間的相關(guān)分析表明(表2),產(chǎn)量與穗粒重、單穗重、出谷率和公頃穗數(shù)呈顯著正相關(guān),與生育期呈顯著負(fù)相關(guān);生育期與穗粒重、出谷率、千粒重、公頃穗數(shù)呈顯著負(fù)相關(guān);說(shuō)明在該區(qū)可以通過(guò)適當(dāng)?shù)目s短生育期,來(lái)提高谷子的結(jié)實(shí)性及成穗數(shù),進(jìn)而達(dá)到提高產(chǎn)量的目的。
表2 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種主要農(nóng)藝性狀及產(chǎn)量平均值間的相關(guān)分析
*<0.05;**<0.01
2.2.4 抗逆性 通過(guò)鑒定的品種均為中抗以上類型,2005—2015年總體抗性有增加的趨勢(shì),其中,品種的抗倒性優(yōu)于對(duì)照;紅葉病和白發(fā)病作為該區(qū)谷子的主要病害在不同年份、不同品種都有發(fā)生,抗白發(fā)病的品種有長(zhǎng)農(nóng)44號(hào)、長(zhǎng)生08、晉谷48、長(zhǎng)農(nóng)39號(hào),抗紅葉病品種有晉谷58;蛀莖率、紋枯病、谷瘟病有升高的趨勢(shì);2個(gè)抗除草劑品種的蛀莖率最高;在對(duì)照倒伏年份里,未發(fā)生倒伏的品種株高都較矮。燕谷16矮稈抗倒、抗白發(fā)病、成穗數(shù)高、但出谷率低。長(zhǎng)生07矮稈、穗長(zhǎng)、生育期長(zhǎng)、穗粒重、單穗重高,但出谷率較低,產(chǎn)量增幅最高,抗性中等以上(電子附表2)。
2.2.5 品種類型及育種方法 2005—2015年通過(guò)鑒定的30個(gè)品種中,除長(zhǎng)生07為系選外,其余品種均為雜交選育的品種,育種方法較單一。品種類型還不夠豐富,30個(gè)品種中僅有抗除草劑品種2個(gè),糧草兼用型品種1個(gè),糯質(zhì)品種1個(gè)。從植株高度看,多數(shù)品種為中稈及以上品種,但與對(duì)照長(zhǎng)農(nóng)35號(hào)相比,通過(guò)鑒定品種的株高有降低趨勢(shì),株高在135 cm以下的中矮稈品種有8個(gè),抗倒性也提高了,但與生產(chǎn)要求還有一定差距,不利于機(jī)械化收獲。穗松緊度適中的品種有22個(gè),穗緊的7個(gè),穗松的1個(gè);穗型為紡錘型的17個(gè),圓筒型8個(gè),棍棒型4個(gè),鞭型1個(gè)。穗長(zhǎng)在17.7—26.2 cm,多為中短穗,穗長(zhǎng)25 cm以上品種2個(gè)。谷粒顏色黃色16個(gè),白色14個(gè);米色均為黃色;熟相中等品種15個(gè),好的15個(gè)??偟膩?lái)說(shuō),春谷中晚熟區(qū)谷子育種手段較單一,品種類型不多,品種多為中高稈品種,矮稈品種選育進(jìn)展較慢,品種穗碼松緊度以中等為主,穗型紡錘形居多,谷穗顏色以黃和白為主,小米顏色均為黃色,熟相中等偏好(電子附表3)。
2.2.6 品種品質(zhì)表現(xiàn) 對(duì)2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種的粗蛋白、粗脂肪、直鏈淀粉、膠稠度、VB1、糊化溫度堿消指數(shù)和賴氨酸含量等品質(zhì)分析結(jié)果(該數(shù)據(jù)由品種選育單位提供,所以數(shù)據(jù)來(lái)源不一致)進(jìn)行了統(tǒng)計(jì)分析,通過(guò)鑒定的品種有29個(gè)進(jìn)行了粗蛋白含量測(cè)定,其中22個(gè)含量分布在10%—14%,2個(gè)低于10%,都為一級(jí)優(yōu)質(zhì)米,5個(gè)高于14%,1個(gè)二級(jí)優(yōu)質(zhì)米;28個(gè)品種進(jìn)行了粗脂肪含量測(cè)定,其中26個(gè)品種含量分布在2%—5%,有2個(gè)低于2%,一、二級(jí)優(yōu)質(zhì)米各1個(gè);25個(gè)品種進(jìn)行了直鏈淀粉測(cè)定,其中18個(gè)品種含量分布在15%—21%,有2個(gè)品種低于15%,其中汾選8號(hào)為0.24%為糯性品種,有5個(gè)品種高于21%;24個(gè)品種進(jìn)行了膠稠度測(cè)定,其中20個(gè)品種分布在115—155 mm內(nèi),2個(gè)品種低于115 mm,一、二級(jí)優(yōu)質(zhì)品種各1個(gè),2個(gè)品種高于155 mm;19個(gè)品種進(jìn)行了VB1的測(cè)定,其中10個(gè)品種分布在3—4 mg·kg-1,有4個(gè)品種低于3 mg·kg-1,有5個(gè)品種高于5 mg·kg-1;24個(gè)品種進(jìn)行了糊化溫度堿消指數(shù)測(cè)定,其中23個(gè)品種分布在3—4,有1個(gè)品種超過(guò)了4,為一級(jí)優(yōu)質(zhì)米;8個(gè)品種進(jìn)行了賴氨酸的測(cè)定,含量分布在0.18%—0.24%,優(yōu)質(zhì)米中含量相對(duì)較高??傊?,優(yōu)質(zhì)米的粗蛋白、粗脂肪含量及膠稠度偏低,賴氨酸含量相對(duì)較高(電子附表4、電子附圖2)。
2005—2015年西北春谷中晚熟區(qū)參試品種及通過(guò)鑒定品種的產(chǎn)量(圖1和圖2)有逐年增加的趨勢(shì),但和對(duì)照相比,參試品種逐年變化趨勢(shì)和對(duì)照接近,通過(guò)鑒定品種增產(chǎn)幅度不大。該區(qū)11年來(lái)育成品種在產(chǎn)量上整體沒(méi)有大的突破,30個(gè)鑒定品種中,除長(zhǎng)生07為系選外,其余品種均為雜交選育的品種,育種手段、品種類型較單一。通過(guò)鑒定品種在產(chǎn)量提高上發(fā)揮的作用不大,單產(chǎn)提高可能是氣候[26-27]或者栽培環(huán)境的影響更大。
西北春谷中晚熟區(qū)的株高(圖2)波動(dòng)在105—165 cm,變幅較大,除少量矮稈品種外,多數(shù)品種為中高稈類型品種,蒙金谷一號(hào)和晉谷55號(hào)的株高超過(guò)162 cm,其抗倒性明顯下降。影響谷子抗倒性的因素很多,矮稈并非抗倒的唯一因素[28-30],已有研究表明,谷子的根量以及莖稈的機(jī)械強(qiáng)度和抗折力對(duì)谷子品種的抗倒伏能力有顯著影響[31],但株高是其中主要因素。培育矮稈品種是提高抗倒性和產(chǎn)量的有效途徑,也符合培育適合機(jī)械化收獲品種的要求。近年來(lái)育成品種的株高呈現(xiàn)出降低的趨勢(shì),是谷子育種新方向。對(duì)通過(guò)鑒定的品種農(nóng)藝性狀和產(chǎn)量的相關(guān)性分析表明,生育期和產(chǎn)量為負(fù)相關(guān)(表2),說(shuō)明生育期過(guò)長(zhǎng)不利于谷子產(chǎn)量的提高,影響了品種的適應(yīng)范圍,不利于應(yīng)對(duì)氣候的變化。近幾年育成品種的生育期呈縮短的趨勢(shì),多數(shù)低于130 d(電子附圖1),在120 d上下波動(dòng),通過(guò)適當(dāng)?shù)目s短生育期,提高谷子的結(jié)實(shí)性及成穗數(shù),進(jìn)而達(dá)到提高產(chǎn)量的目的。種植密度近年來(lái)呈現(xiàn)增大趨勢(shì)(33萬(wàn)—43萬(wàn)株/hm2),提高谷子的產(chǎn)量不僅要提高品種的豐產(chǎn)性,還應(yīng)通過(guò)良種良法配套實(shí)現(xiàn)高產(chǎn)??偟膩?lái)說(shuō),春谷中晚熟區(qū)品種的選育應(yīng)重點(diǎn)考慮降低株高、縮短生育期、提高穗重、改善結(jié)實(shí)性、增加成穗數(shù),在提高產(chǎn)量的同時(shí),適應(yīng)谷子機(jī)械化生產(chǎn)的需求。
2005—2015年通過(guò)鑒定品種品質(zhì)特性(電子附表3、電子附圖2)中,米色全為黃色,優(yōu)質(zhì)米的數(shù)量超過(guò)了50%。品質(zhì)結(jié)果分析表明,優(yōu)質(zhì)米的粗蛋白、粗脂肪含量及膠稠度偏低,賴氨酸含量相對(duì)較高??傮w上看,品種的品質(zhì)取得較大進(jìn)步,同時(shí)包括糯性品種(直鏈淀粉含量?jī)H為0.24%)、高蛋白品種(粗蛋白含量高于14%)、低糊化溫度品種(糊化溫度堿消指數(shù)高于4)等多種類型,品種的多樣性對(duì)于市場(chǎng)多樣化需求具有重要的意義。彭鎖堂等[32]對(duì)四大名米之一“沁州黃小米”的糊化溫度、膠稠度、直鏈淀粉含量進(jìn)行測(cè)定表明,沁州黃小米優(yōu)質(zhì)的原因在于具有中等偏低的直鏈淀粉含量和糊化溫度,以及中等偏高的米膠長(zhǎng)度。從市場(chǎng)需求看,市場(chǎng)對(duì)優(yōu)質(zhì)米的要求一般主要看小米的色澤、一致性以及適口性,但國(guó)內(nèi)外對(duì)谷子優(yōu)質(zhì)品種的評(píng)價(jià)沒(méi)有一個(gè)能夠排除人為因素的量化方法。中國(guó)作物學(xué)會(huì)粟類專業(yè)委員會(huì)組織的“全國(guó)優(yōu)質(zhì)食用粟品質(zhì)鑒評(píng)”,采取百分制人為打分的方法,評(píng)選結(jié)果能夠更接近市場(chǎng)的需求,品質(zhì)化驗(yàn)結(jié)果在一定程度上反映了品種的品質(zhì),特別是適口性相關(guān)指標(biāo),對(duì)于指導(dǎo)優(yōu)質(zhì)品種選育具有很好的指導(dǎo)作用。
2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定的品種多達(dá)30個(gè),數(shù)量上不少,其中有飼草品種1個(gè),糯性品種1個(gè),抗除草劑品種2個(gè),這些特異性品種的育成,表明在谷子資源創(chuàng)新、簡(jiǎn)化育種和谷子多元化育種方面取得了一定的進(jìn)展,對(duì)推動(dòng)谷子產(chǎn)業(yè)發(fā)展發(fā)揮了積極的作用。從育種方法上看,西北春谷區(qū)育成的品種多采用簡(jiǎn)單的雜交、系選等方法,根據(jù)育種目標(biāo)采取回交、復(fù)交以及理化誘變等方法上應(yīng)用的少,雖然取得了一定的效果,但由于應(yīng)用方法自身的特點(diǎn)和人為對(duì)少數(shù)骨干親本的集中利用,造成育成品種遺傳基礎(chǔ)狹窄[33-34],品種的產(chǎn)量、抗性和品質(zhì)等方面均不能滿足市場(chǎng)多樣性需求及谷子產(chǎn)業(yè)化生產(chǎn)發(fā)展的需要。針對(duì)市場(chǎng)需求和谷子產(chǎn)業(yè)化發(fā)展,挖掘、利用和創(chuàng)新優(yōu)質(zhì)特異谷子種質(zhì)資源,建立目標(biāo)性狀基因庫(kù)[35-38],結(jié)合分子生物學(xué)技術(shù)和克隆基因技術(shù),建立現(xiàn)代生物技術(shù)范疇的基因庫(kù),將常規(guī)育種與分子育種新技術(shù)結(jié)合,開(kāi)展多元化、多目標(biāo)育種,培育性狀優(yōu)良、抗性強(qiáng)、適應(yīng)性廣、產(chǎn)量高、品質(zhì)優(yōu)良、適合機(jī)械化生產(chǎn)的谷子新品種是該區(qū)域谷子育種未來(lái)發(fā)展方向。
西北春谷晚熟區(qū)谷子品種的選育取得了一定的進(jìn)步,所育品種在數(shù)量、多樣性、產(chǎn)量、品質(zhì)等方面有了一定的提高,但沒(méi)有取得大的突破。在育種方法上比較單一,多采取簡(jiǎn)單雜交、系選的方法,應(yīng)根據(jù)育種目標(biāo)要求,開(kāi)展材料創(chuàng)新,融合回交、復(fù)交、理化誘變等多種方法,并結(jié)合分子育種新技術(shù),提高育種水平。在育種目標(biāo)上,重點(diǎn)培育品質(zhì)優(yōu)良、矮稈抗倒、生育期略短、適合機(jī)械化收獲的品種,培育多種類型的抗除草劑品種,滿足谷子規(guī)?;a(chǎn)的需求。
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(責(zé)任編輯 李莉)
A:生育期;B:株高;C:穗長(zhǎng);D:?jiǎn)嗡胫兀籈:穗粒重;F:千粒重;G:出谷率;H:公頃穗數(shù)A: Growth duration; B: Plant height; C: Panicle length; D: Weight per panicle; E: Grain weight per panicle; F: 1000-grain weight; G: Percentage of grain weight per panicle; H: Number of panicles per hectare
1:長(zhǎng)0301 Chang 0301;2:長(zhǎng)農(nóng)36號(hào)Changnong 36;3:汾選4號(hào)Fenxuan 4;4:太選4號(hào);Taixuan 4;5:汾選8號(hào)Fenxuan 8;6:蒙谷10號(hào) Menggu 10;7:晉谷42號(hào)Jingu 42;8:晉谷45號(hào)Jingu 45;9:長(zhǎng)農(nóng)38號(hào)Changnong 38;10:長(zhǎng)生07 Changsheng 07;11:朝谷15號(hào)Chaogu 15; 12:晉谷48號(hào)Jingu 48;13:燕谷16 Yangu 16;14:興谷66Xinggu 66;15:蒙金谷一號(hào)Mengjingu 1;16:晉谷51號(hào)Jingu 51;17:長(zhǎng)農(nóng)39號(hào) Changnong 39;18:長(zhǎng)生08 Changsheng 08;19:承谷13Chenggu 13;20:長(zhǎng)生10號(hào)Changsheng 10;21:晉汾02 Jinfen 02;22:晉谷55號(hào)Jingu 55;23:豫谷18 Yugu 18;24:晉谷58 Jingu 58;25:朝谷19 Chaogu 19;26:長(zhǎng)農(nóng)44號(hào)Changnong 44;27:長(zhǎng)生11 Changsheng 11;28:晉谷59號(hào) Jingu 59;29:晉谷62號(hào)Jingu 62;30:朝谷58 Chaogu 58
附圖1 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種與對(duì)照品種農(nóng)藝性狀平均值的比較
Fig. S1 The agronomic traits of cultivars released compared with CK
附圖2 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種品質(zhì)性狀
Fig. S2 The quality characters of cultivars released in 2005-2015
附表1 2005-2015年西北春谷中晚熟區(qū)通過(guò)鑒定的品種
Table S1 The foxtail millet cultivars released in the regional adaptation tests in 2005-2015
編號(hào)Number品種名稱Cultivar來(lái)源Sources選育單位Breeding institution特異性狀Specific traits優(yōu)質(zhì)米等級(jí)Quality level鑒定編號(hào)GIA report number 1長(zhǎng)0301Chang 0301[(683×?xí)x20)長(zhǎng)20] ×98-12[(683×Jin 20)Chang 20] ×98-12山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences2007006 2長(zhǎng)農(nóng)36號(hào)Changnong 36長(zhǎng)農(nóng)28 ×92新矮Changnong 28 ×92 Xinai山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences一級(jí)The first quality level 2007008 3汾選4號(hào)Fenxuan 43173×?xí)x谷21號(hào)3173×Jingu 21山西省農(nóng)業(yè)科學(xué)院經(jīng)濟(jì)作物研究所Economic Crops Research Institute of Shanxi Academy of Agricultural Sciences2007009 4太選4號(hào)Taixuan 495品10×?xí)x谷20號(hào)95 Pin 10×Jingu 20山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences2007007 5汾選8號(hào)Fenxuan 88610×?xí)x谷21號(hào)8610×Jingu 21山西省農(nóng)業(yè)科學(xué)院經(jīng)濟(jì)作物研究所Economic Crops Research Institute of Shanxi Academy of Agricultural Sciences糯質(zhì)品種Waxy variety2009015 6蒙谷10號(hào)Menggu 10有性雜交選育(草谷系選)Breeding by sexual hybridization(selected from Caogu)內(nèi)蒙古農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of inner Mongolia Academy of Agricultural Sciences二級(jí)The second quality level2008006 7晉谷42號(hào)Jingu 42960011×?xí)x谷21號(hào)960011×Jingu 21山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences2008007 8晉谷45號(hào)Jingu 45赤谷8號(hào)×?xí)x谷20號(hào)Chigu 8×Jingu 20山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences2009005 9長(zhǎng)農(nóng)38號(hào)Changnong 3896r-20×91品896r-20×91Pin 8山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences2009004 10長(zhǎng)生07Changsheng 07長(zhǎng)農(nóng)35號(hào)系選Selected from Changnong 35山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences二級(jí)The second quality level2009003 續(xù)附表1 Continued table S1 編號(hào)Number品種名稱Cultivar來(lái)源Sources選育單位Breeding institution特異性狀Specific traits優(yōu)質(zhì)米等級(jí)Quality level鑒定編號(hào)GIA report number 11朝谷15號(hào)Chaogu 15鐵谷7號(hào)×朝谷9號(hào)Tiegu 7×Chaogu 9遼寧省農(nóng)業(yè)科學(xué)院水土保持研究所Soil and Water Conservation Institute of Liaoning Academy of Agricultural Sciences二級(jí)The second quality level2009014 12晉谷48號(hào)Jingu 48晉谷30號(hào)×94-158Jingu 30 ×94-158山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences2009011 13燕谷16Yangu 16朝谷13×昭農(nóng)21Chaogu 13×Zaonong 21遼寧省農(nóng)業(yè)科學(xué)院水土保持研究所Soil and Water Conservation Institute of Liaoning Academy of Agricultural Sciences二級(jí)The second quality level2009012 14興谷66Xinggu 66束離子處理晉谷28號(hào)山西省農(nóng)業(yè)科學(xué)院Shanxi Academy of Agricultural Sciences2009013 15蒙金谷一號(hào)Mengjingu 1朱砂谷×品系83574Zhushagu×Pinxi 83574內(nèi)蒙古農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Inner Mongolia Academy of Agricultural Sciences糧飼兼用品種Food and Feed variety一級(jí)The first quality level2010010 16晉谷51號(hào)Jingu 51晉谷30號(hào)×品谷2號(hào)晉谷30號(hào)×品谷2號(hào)Jingu 30×Pingu 2山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences2010004 17長(zhǎng)農(nóng)39號(hào)Changnong 3997F高-66×?xí)x谷21號(hào)97F Gao-66×Jingu 21山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences二級(jí)The second quality level2010005 18長(zhǎng)生08Changsheng 08長(zhǎng)204×?xí)x谷21號(hào)Chang 204×Jin gu 21山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences2010006 19承谷13Chenggu 13豫谷2×承農(nóng)2Yugu 2×Chengnong 2河北省承德市農(nóng)業(yè)科學(xué)研究所Chengde Academy of Agricultural Sciences in Hebei Province二級(jí)The second quality level2010007 20長(zhǎng)生10號(hào)Changsheng 10(GM×矮88)A×混合父本(GM×Ai 88)A×Mixed parents山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences二級(jí)The second quality level201308 21晉汾02Jinfen 0287-160×(148×8531)山西省農(nóng)業(yè)科學(xué)院經(jīng)濟(jì)作物研究所Economic Crops Research Instituteof Shanxi Academy of Agricultural Sciences一級(jí)The first quality level201310 續(xù)附表1 Continued table S1 編號(hào)Number品種名稱Cultivar來(lái)源Sources選育單位Breeding institution特異性狀Specific traits優(yōu)質(zhì)米等級(jí)Quality level鑒定編號(hào)GIA report number 22晉谷55號(hào)Jingu 55970004×?xí)x谷36號(hào)970004×Jingu 36山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences201309 23豫谷18Yugu 18豫谷1號(hào)×保282Yugu 1×Bao 282河南省安陽(yáng)市農(nóng)業(yè)科學(xué)院Anyang Academy of Agricultural Sciences in Henan Province一級(jí)The first quality level2014009 24晉谷58Jingu 58晉谷29號(hào)×赤谷8號(hào)Jin gu 29×Chigu 8山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences2014011 25朝谷19Chaogu 19朝谷13自然雜交株Nature crossing from Chaogu 13 遼寧省農(nóng)業(yè)科學(xué)院水土保持研究所Soil and Water Conservation Institute of Liaoning Academy of Agricultural Sciences二級(jí)The second quality level2014010 26長(zhǎng)農(nóng)44號(hào)Changnong 442007品11×RN2007Pin 11/×RN山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of shanxi Academy of Agricultural Sciences抗拿捕凈Resist sethoxydim二級(jí)The second quality level2015011 27長(zhǎng)生11Changsheng 11長(zhǎng)農(nóng)35系選Selected from Changnong 35山西省農(nóng)業(yè)科學(xué)院谷子研究所Millet Research Institute of Shanxi Academy of Agricultural Sciences2015010 28晉谷59號(hào)Jingu 59晉谷30號(hào)×?xí)x谷36號(hào)Jingu 30×Jingu 36山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences一級(jí)The first quality level2015009 29晉谷62號(hào)Jingu 6202-68×?xí)x谷36號(hào)02-68×Jingu 36山西省農(nóng)業(yè)科學(xué)院作物科學(xué)研究所Crop Sciences Research Institute of Shanxi Academy of Agricultural Sciences二級(jí)The second quality level2016013 30朝谷58Chaogu 58朝谷9號(hào)×冀谷25Chaogu 9×Jigu 25遼寧省農(nóng)業(yè)科學(xué)院水土保持研究所Soil and Water Conservation Institute of Liaoning Academy of Agricultural Sciences抗拿捕凈Resist sethoxydim二級(jí)The second quality level2016012
附表2 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種抗逆性
Table S2 The resistance of foxtail millet cultivars released in regional adaptation test in 2005-2015
區(qū)試時(shí)間RAT 序號(hào)Number品種名稱Cultivar抗倒性Lodging(0-4 level)耐旱性Drought tolerance (0-4 level)谷銹病Rust-resistance(0-4 level)谷瘟病Blast-resistance (0-4 level)紋枯病Sheath blight- resistance(0-4 level)紅葉病The rate of red leaf disease (%)白發(fā)病The rate of downy- mildew (%)黑穗病The rate of grain smut (%)線蟲(chóng)病The rate of worm disease (%)蛀莖率The rate of bethyloid(%) 2005-2006 1長(zhǎng)0301Chang 03011321171.5003.5 2長(zhǎng)農(nóng)36號(hào)Changnong 362331162005 3汾選4號(hào)Fenxuan 42232186003.5 4太選4號(hào)Taixuan 4033225.53.2004 CK承谷8號(hào)Chenggu 82231091.5003 2006-2007 5汾選8號(hào)Fenxuan 832312108.7005 6蒙谷10號(hào)Menggu 102242284.2000.05 7晉谷42號(hào)Jingu 42132116.54.59.600.03 CK2006承谷8號(hào)Chenggu 82221091.5000.03 CK2007長(zhǎng)農(nóng)35號(hào)Changnong 35012111.86.2000 2007-2008 8晉谷45號(hào)Jingu 451131233.2000 9長(zhǎng)農(nóng)38號(hào)Changnong 3821211512.1003 10長(zhǎng)生07Changsheng 071131139002 11朝谷15號(hào)Chaogu 1502111103.3001 CK長(zhǎng)農(nóng)35號(hào)Changnong 352121136.2004 2008-200912晉谷48號(hào)Jingu 484210140.6002.3 13燕谷16Yangu 1602111151.101.392.7 14興谷66Xinggu 660211134.49002.7 15蒙金谷一號(hào)Mengjingu 142211101001.7 CK長(zhǎng)農(nóng)35號(hào)Changnong 352221135.66004 2009-201016晉谷51號(hào)Jingu 51121011.221.4004 17長(zhǎng)農(nóng)39號(hào)Changnong 39121113.90.9001 18長(zhǎng)生08Changsheng 081121210.4002 19承谷13Chenggu 134211116.251.1002.5 續(xù)附表2 Continued table S2 區(qū)試時(shí)間RAT 序號(hào)Number品種名稱Cultivar抗倒性Lodging(0-4 level)耐旱性Drought tolerance (0-4 level)谷銹病Rust-resistance(0-4 level)谷瘟病Blast-resistance (0-4 level)紋枯病Sheath blight- resistance(0-4 level)紅葉病The rate of red leaf disease (%)白發(fā)病The rate of downy- mildew (%)黑穗病The rate of grain smut (%)線蟲(chóng)病The rate of worm disease (%)蛀莖率The rate of bethyloid(%) CK長(zhǎng)農(nóng)35號(hào)Changnong 350211109.57001 2010-201120長(zhǎng)生10號(hào)Changsheng 101121132.56002.6 CK長(zhǎng)農(nóng)35號(hào)Changnong 35301119.30.5000.4 2011-201221晉汾02Jinfen 023012110.615.5002.8 22晉谷55號(hào)Jingu 55322113.30.88005.2 CK長(zhǎng)農(nóng)35號(hào)Changnong 35322119.31.38005.2 2012-201323豫谷18Yugu 18223338.43.87006.04 24晉谷58Jingu 58223330.41.10011.01 25朝谷19Chaogu 19231336.51001.89 CK長(zhǎng)農(nóng)35號(hào)Changnong 35223331.21.4008.89 2013-201426長(zhǎng)農(nóng)44號(hào)Changnong 442233350.80020.12 27長(zhǎng)生11Changsheng 111231334.4005.83 28晉谷59號(hào)Jingu 59123331.81.10015.02 CK長(zhǎng)農(nóng)35號(hào)Changnong 353233324.1008.89 2014-201529晉谷62號(hào)Jingu 62121112.211.22005.33 30朝谷58Chaogu 58022011.99.940016.93 CK長(zhǎng)農(nóng)35號(hào)Changnong 353213224.950011.23
粗體為不同年度對(duì)照的抗逆性
The bold letters are the resistance of CK in different year; RAT: Regional adaptation test
附表3 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種株形及穗部性狀
Table S3 The panicle characters of foxtail millet cultivars released in the regional adaptation test in 2005-2015
序號(hào)Number品種名稱Cultivar株莖高Plant height (cm)穗長(zhǎng)Panicle length (cm)穗松緊Density of panicle穗型Type of panicle谷色Grain color米色Millet color熟相Maturity of plant 1長(zhǎng)0301Chang 0301139.518.9中higher棍棒Club黃Yellow黃Yellow好Best 2長(zhǎng)農(nóng)36號(hào)Changnong 36142.720.5緊highest紡綞Spindle白White黃Yellow好Best 3汾選4號(hào)Fenxuan 4138.921.8中higher紡綞Spindle黃Yellow黃Yellow中Better 4太選4號(hào)Taixuan 4130.922.0中higher圓筒Cylindrical黃Yellow黃Yellow好Best 5汾選8號(hào)Fenxuan 8133.523.1松lower紡綞Spindle黃Yellow黃Yellow中Better 6蒙谷10號(hào)Menggu 10144.526.2中higher紡綞Spindle白White黃Yellow中Better 7晉谷42號(hào)Jingu 42131.420.6中higher圓筒Cylindrical黃Yellow黃Yellow中Better 8晉谷45號(hào)Jingu 45155.321.4中higher紡綞Spindle白White黃Yellow中Better 9長(zhǎng)農(nóng)38號(hào)Changnong 38139.921.4中higher紡綞Spindle白White黃Yellow中Better 10長(zhǎng)生07Changsheng 07135.920.4緊highest紡綞Spindle白White黃Yellow中Better 11朝谷15號(hào)Chaogu 15120.723.0中higher紡綞Spindle白White黃Yellow中Better 12晉谷48號(hào)Jingu 48134.619.8中Higher棍棒Club黃Yellow黃Yellow好Best 13燕谷16Yangu 16105.522.0中Higher圓筒Cylindrical白White黃Yellow中Better 14興谷66Xinggu 66140.123.9緊Highest鞭Whip白White黃Yellow中Better 15蒙金谷一號(hào)Mengjingu 1162.121.7中Higher紡綞Spindle白White黃Yellow中Better 16晉谷51號(hào)Jingu 51141.221.1緊Highest棍棒Club黃Yellow黃Yellow好Best 17長(zhǎng)農(nóng)39號(hào)Changnong 39135.923.3中Higher紡綞Spindle白White黃Yellow好Best 18長(zhǎng)生08Changsheng 08146.622.7中Higher紡綞Spindle黃Yellow黃Yellow好Best 19承谷13Chenggu 13130.921.7緊Highest紡綞Spindle白White黃Yellow好Best 20長(zhǎng)生10號(hào)Changsheng 10158.617.7中Higher圓筒Cylindrical白White黃Yellow中Better 21晉汾02Jinfen 02138.420.0中Higher紡綞Spindle白White黃Yellow好Best 22晉谷55號(hào)Jingu 55163.025.0中Higher紡綞Spindle黃Yellow黃Yellow中Better 23豫谷18Yugu 18120.722.7中Higher圓筒Cylindrical黃Yellow黃Yellow好Best 24晉谷58Jingu 58144.222.4緊Highest圓筒Cylindrical黃Yellow黃Yellow中Better 25朝谷19Chaogu 19155.124.4中Higher紡綞Spindle黃Yellow黃Yellow中Better 26長(zhǎng)農(nóng)44號(hào)Changnong 44137.223.7中Higher紡綞Spindle黃Yellow黃Yellow好Best 27長(zhǎng)生11Changsheng 11147.319.3緊Highest棍棒Club白White黃Yellow好Best 28晉谷59號(hào)Jingu 59153.421.6中Higher圓筒Cylindrical黃Yellow黃Yellow好Best 29晉谷62號(hào)Jingu 62147.524.0中Higher圓筒Cylindrical黃Yellow黃Yellow好Best 30朝谷58Chaogu 58124.420.8中Higher紡綞Spindle黃Yellow黃Yellow好Best
附表4 2005—2015年西北春谷中晚熟區(qū)通過(guò)鑒定品種品質(zhì)性狀
Table S4 The quality characters of cultivars released in the regional adaptation test in 2005-2015
序號(hào)Number粗蛋白含量Crude protein content(%)粗脂肪含量Crude fat content (%)VB1(mg·kg-1)直鏈淀粉Amylose content(%)膠稠度Gel consistency(mm)堿消指數(shù)Alkali index elimination賴氨酸Lysine備注Note 長(zhǎng)農(nóng)35號(hào)Changnon 3513.13.629.214.181052.80.31一級(jí)優(yōu)質(zhì)米 The first quality level 212.773.9117.561154.70.24一級(jí)優(yōu)質(zhì)米 The first quality level 158.83一級(jí)優(yōu)質(zhì)米 The first quality level 2113.572.412.4116.53.80.24一級(jí)優(yōu)質(zhì)米 The first quality level 238.941.58一級(jí)優(yōu)質(zhì)米 The first quality level 2812.913.133.115.9131.53.6一級(jí)優(yōu)質(zhì)米 The first quality level 614.451.272.618.731034二級(jí)優(yōu)質(zhì)米 The second quality level 1011.652.423.716.75136.53.70.21二級(jí)優(yōu)質(zhì)米 The second quality level 1110.392.5922.81833.70.18二級(jí)優(yōu)質(zhì)米 The second quality level 1310.63二級(jí)優(yōu)質(zhì)米 The second quality level 1712.723.545.515.531453.2二級(jí)優(yōu)質(zhì)米 The second quality level 2011.643.0118.691294二級(jí)優(yōu)質(zhì)米 The second quality level 2510.552.1426.750.18二級(jí)優(yōu)質(zhì)米 The second quality level 2610.493.384.216.07125.53.6二級(jí)優(yōu)質(zhì)米 The second quality level 2911.834.075.622.14117.53二級(jí)優(yōu)質(zhì)米 The second quality level 3011.262.1123.620.18二級(jí)優(yōu)質(zhì)米 The second quality level 112.782.853.215.561483.60.21 314.483.5615.961364 414.594.435.417.17117.53.7 511.474.050.241884 711.934.3618.7117.53.8 815.474.544.816.24132.53.7 912.32.172.318.31603.90.18 1210.883.844.518.431303.1 14 1611.553.815.218.131353.2 1812.883.772.920.831313.7 1913.614.374.315.8132.53.8 2212.084.034.915.41132.53.8 2411.474.173.721.391183.7 2715.634.374.913.79136.53.6
Evaluation of Foxtail Millet Cultivars Developed in the Middle and late-Maturing Spring-sowing Region in Northwest China in 2005-2015
ZHANG AiYing1, GUO ErHu1, DIAO XianMin2, FAN HuiPing1, LI YuHui1, WANG LiXia1, GUO HongLiang1, CHENG LiPing1, Wu YinSheng1
(1Millet Research Institute, Shanxi Academy of Agricultural Sciences/Shanxi Key Laboratory of Genetic Resources and Breeding in Minor Crops, Changzhi 046011, Shanxi;2Institute of Crop Science, Chinese Academy of Agricultural Sciences, Beijing 100081)
The objective of this study is to analyze the changes of agronomic traits, grain yield, stress resistance and quality traits of newly developed foxtail millet cultivars in the middle and late-maturing spring-sowing region in northwest China in 2005-2015-, and provide scientific data for new cultivar development and genetic resource enhancement.Based on the data of middle and late-maturing spring-sowing foxtail millet lines tested in the regional adaptation tests from 2005 to 2015 in northwest China, main agronomic traits and yield traits were analyzed and compared. Diseases resistances and quality traits of 30 foxtail millet cultivars released were further analyzed in this paper.The grain yield potential of the newly developed foxtail millet cultivars in 2005-2015 showed a trend of increase from the data analyzed. Great difference of agronomic characteristics of tested lines and released cultivars among experimental years were identified, which implied that climate difference among years might have great impact on foxtail millet characters. The data showed a clear increasing trend in characteristics such as plant height, panicle weight, panicle grain weight and ratio of grain on a panicle of the newly developed cultivars, while the growth duration of those varieties became shorter than before. Coefficient correlation analyses of the agronomic traits indicated that grain yield of cultivars is positively correlated with panicle weight, grain weight per panicle and number of panicles per hectare and negatively related with the length of growth duration. The commonly occurred diseases in this foxtail millet growing region included leaf rust, blast, sheath blight, red leaf disease and downy-mildew in the past ten years and among which the latter two are the main ones. All the newly developed cultivars showed their increase in the abilities of diseases resistance. Most newly developed cultivars were characterized with yellow kernel, and the most adapted cultivars were usually characterized with a relatively lower content of crude protein and fatty acids, but a relatively higher lysine content. Different types of cultivars including waxy starch, high protein content, herbicide resistant were developed in the past ten years which provided firm foundation for foxtail millet industry in this region.The yield potential of newly developed foxtail millet cultivars in the middle and late maturing region of northwest China in the past 11 years was improved. Different types of cultivars were developed so as to meet foxtail millet industry in this region. The main method of foxtail millet breeding in this region is pedigree selection after hybridization between cultivars, which should be improved with other methods such as molecular marker assistant selection.
spring foxtail millet; middle and late maturing region; regional adaptation test; agronomic characters; correlation analysis
2017-05-12;
2017-06-16
國(guó)家現(xiàn)代農(nóng)業(yè)產(chǎn)業(yè)技術(shù)體系建設(shè)專項(xiàng)(CARS-07-13.5-A10)、山西省農(nóng)業(yè)科學(xué)院生物育種工程項(xiàng)目(17yzgc024)、山西省農(nóng)業(yè)科學(xué)院農(nóng)業(yè)科技創(chuàng)新研究課題(YYS1703)
聯(lián)系方式:張艾英,E-mail:zay1012@126.com。通信作者郭二虎,E-mail:guoerhu2003@163.com。通信作者刁現(xiàn)民,E-mail:diaoxianmin@caas.cn