• <tr id="yyy80"></tr>
  • <sup id="yyy80"></sup>
  • <tfoot id="yyy80"><noscript id="yyy80"></noscript></tfoot>
  • 99热精品在线国产_美女午夜性视频免费_国产精品国产高清国产av_av欧美777_自拍偷自拍亚洲精品老妇_亚洲熟女精品中文字幕_www日本黄色视频网_国产精品野战在线观看 ?

    Restudy of the Late Oligocene dormice from northern Junggar Basin

    2016-03-29 03:07:19WUWenYuMENGJin2YEJieNIXiJunBIShunDong
    古脊椎動物學報(中英文) 2016年1期
    關鍵詞:印第安納哈巴準噶爾盆地

    WU Wen-YuMENG Jin,2YE JieNI Xi-JunBI Shun-Dong,3

    (1Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences,Institute of Vertebrate Paleontology and Paleoanthropology,Chinese Academy of SciencesBeijing 100044, China wuwenyu@ivpp.ac.cn)

    (2Division of Paleontology,American Museum of Natural HistoryNew York NY 10024, USA)

    (3Department of Biology,Indiana University of PennsylvaniaIndiana PA 15705, USA)

    Restudy of the Late Oligocene dormice from northern Junggar Basin

    WU Wen-Yu1MENG Jin1,2YE Jie1NI Xi-Jun1BI Shun-Dong1,3

    (1Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences,Institute of Vertebrate Paleontology and Paleoanthropology,Chinese Academy of SciencesBeijing 100044, China wuwenyu@ivpp.ac.cn)

    (2Division of Paleontology,American Museum of Natural HistoryNew York NY 10024, USA)

    (3Department of Biology,Indiana University of PennsylvaniaIndiana PA 15705, USA)

    A new glirid genus and species,Gliruloides zhoui, is named based on specimens from the Late Oligocene Tieersihabahe Mammal Assemblage Zone I (Tie-I zone) of the northern Junggar Basin, Xinjiang. The new genus is diagnosed by the following features: middle-sized dormouse; occlusal surface of cheek teeth concave; upper and lower cheek teeth dominantly with nine transverse ridges; the anterotrope(id) and posterotrope(id) well developed and extending almost full length of corresponding valleys; transverse ridges of upper cheek teeth usually free-ended labially; M1/2 with V- or narrow U-shaped trigon; the precentroloph not connected to the endoloph that is incomplete or nearly complete; the endolophid in lower cheek teeth discontinuous or continuous; labial end of the anterolophid curving slightly distally but not connected with the protoconid; root number of p4, m1-3, P4 and M1-3 being 2, 2, 3 and 3 respectively. We discuss the differences ofGliruloidesfromGlirulusandVasseuromysand assign the AnatolianVasseuromys duplexandVasseuromysaff.V. duplexfrom the Early Miocene of Turkey toGliruloides. It is posited thatGliruloidesandGlirulusmay share a common ancestor similar toGlisguerbuezifrom the Lower Oligocene of Thrace, Turkey.Gliruloidesmight live in a relative wet and warm biotope.

    northern Junggar Basin, Late Oligocene, Tieersihabahe Mammal Assemblage Zone I, dormouse

    Our study on earliest Chinese dormice (Wu et al., 2000) was based on specimens collected in 1998 from the Late Oligocene Tieersihabahe Formation in northern Junggar Basin of Xinjiang. Only four teeth, among which three were assigned toGlirulussp., were available at the time. Additional 11 teeth were collected by screenwashing at the same level of thesame locality (XJ 98024) and other nearby localities (XJ 98035, XJ 200209 and XJ 20004) in four fi eld seasons from 1999 to 2002. All teeth but one right M3 (IVPP V 18113) are almost identical to the specimens described by Wu et al. (2000). Undoubtedly, they belong to the same taxon. Since the work of Wu et al. (2000), we have realized that these teeth show high similarities toVasseuromysduplexfrom the Early Miocene of Anatolia (ünay, 1994), and therefore citedVasseuromyssp., instead ofGlirulussp., in fauna lists of our successive papers (Meng et al., 2001, 2006; Ye et al., 2001a,b; 2003a,b) but without further explanation. The present paper restudies the Late Oligocene Junggar dormouse based on all material available to us and reinterprets its taxonomic position.

    Our study reveals that the specimens of the Late Oligocene Junggar dormouse are more similar toVasseuromysduplexthan to any other dormouse and thatV. duplexdiffers considerably from other EuropeanVasseuromysspecies andGlirulus. Thus, we name a new genus,Gliruloides, for the Junggar dormouse and assign the Anatolian species,Vasseuromys duplexandVasseuromysaff.V. duplex, to this genus.

    In describing the tooth morphology, we use a modified terminology derived from de Bruijn (1966) and Freudenthal (2004), which is convenient and unambiguous for description of the extra ridges of the dormice cheek teeth (Fig. 1). The SEM photographs were taken by Meng Jin, using the Hitach S-4700 scanning electron microscope at the American Museum of Natural History in 2005.

    Fig. 1 Dental terminology of Gliridae Modi fi ed from de Bruijn (1966) and Freudenthal (2004)

    Family Gliridae Thomas, 1897

    Subfamily Dryomyinae de Bruijn, 1967

    GenusGliruloidesgen. nov.

    Etymologyglirulus+ oides,indicates that the dormouse is morphologically similar to genusGlirulusbut should not beGlirulus.

    Type speciesGliruloides zhouigen. et sp. nov.

    Diagnosis Medium-sized dormouse with concave occlusal surface of the cheek teeth. The upper and lower cheek teeth possess dominantly nine transversal ridges. The anterotrope(id) and posterotrope(id) of the upper and lower cheek teeth are well developed and extend almost full length of the corresponding positioned valleys. The trigon of M1-2 is V-or narrow U-shaped, and the endoloph of upper cheek teeth is incomplete or nearly complete. The main ridges of the upper cheek teeth are labially free-ended. The endolophid of the lower cheek teeth is discontinous or continuous. The labial end of the anterolophid of the lower molars slightly curves distally, but not connected to the protoconid. Lower cheek teeth are two-rooted.

    Differentiate diagnoses 1)Gliruloidesdiffers fromGlirulusin having a V-shaped or narrow U-shaped trigon on M1-2, the endoloph incomplete or nearly complete on upper cheek teeth, the lingual end of the precentroloph not connected to the endoloph, and main ridges in the upper cheek teeth free-ended labially. In contrast, the upper molars ofGlirulushave wide U-shaped trigon, complete endoloph, and the precentroloph that connects lingually to the endoloph (except for geologically older representatives in which the endoloph may be incomplete and the precentroloph unconnected to the endoloph, refer ünay, 1994). InGlirulusthe main tooth ridges usually connect with each other labially and form the anterior and posterior loops (see Fig. 4D).

    2)Gliruloidesdiffers fromVasseuromysmainly in having nine ridges on most upper and lower molars, in having the developed anterotrope and posterotrope on the upper cheek teeth, and the usually regular and continuous extra ridges. InVasseuromys, however, the anterotrope and posterotrope outside the trigon of the upper cheek teeth are absent or weak, and the extra ridges of the cheek teeth are usually irregular and interrupted (see Fig. 4B).

    Included speciesGliruloides duplex(ünay, 1994) from Early Miocene (MN2) of Anatolia, localities Harami 1-3, Turkey.Gliruloidesaff.G. duplex(ünay, 1994) from Early Miocene (MN1) of Anatolia, localities Kil?ak 0, Kil?ak 0" and Kil?ak 3A-3B.

    Gliruloides zhouigen. et sp. nov.

    Glirulussp., Wu et al., 2000

    Vasseuromyssp., Ye et al., 2001a, b, 2003a, b; Meng et al., 2001, 2006

    Etymology In honor of the late Dr. Zhou Mingzhen (Minchen Chow), an academician of the Chinese Academy of Sciences and the pioneer paleomammalogist in China.

    Holotype Left M2, IVPP V 18110.1 (Fig. 2C).

    Paratype One left M3 (IVPP V 18110.2), one right m1 and one left m1 (V 18110.3-4), and one right m3 (V 18110.5) (Fig. 2D, H, J, K).

    Type locality and horizon Locality XJ 98035, Tieersihabahe of northern Junggar Basin, Xinjiang; Tieersihabahe Mammal Assemblage Zone I; Late Oligocene.

    Referred specimens One left P4 (IVPP V 18111.1, Fig. 2A), two right p4 (V 18111.2-3, Fig. 2F-G), one right m1 (V 18111.4, Fig. 2I) from XJ 98024. One left M1/2 (V 18112, Fig.2B) from XJ 200209.

    The specimens described in 2000 include one right M2 (IVPP V 11812.1), one left M3 (V 11812.2) and one left m1 (V 11812.3) from XJ 98024 (Wu et al. 2000: pl. 1, fi gs. 1-3, 5).

    All specimens listed above were from Tieersihabahe of northern Junggar Basin;Tieersihabahe Mammal Assemblage Zone I; Late Oligocene.

    One right M3 (V 18113, Fig. 2E), collected from XJ 20004 at Saerduoyila of northern Junggar Basin. The stratum producing the specimen is correlative to that of Tieersihabahe.

    Diagnoses The p4 possesses eight transverse ridges. The anterotropid on the lower cheek teeth is usually single. The endoloph on upper cheek teeth is nearly complete, whereas the endolophid of lower cheek teeth is interrupted. The protoconid and mesoconid of the lower molars are hook-like. The metalophid extends slightly distally to the labial side and then turns abruptly mesiolabially.

    Differentiate diagnosesGliruloides zhouidiffers from the Anatolian speciesG. duplexandGliruloidesaff.G. duplexin having 1) nearly complete endoloph on M1/2, 2) hook-like protoconid and mesoconid of the lower molars, 3) the metalophid almost always extends slightly distolabially and then turns abruptly mesiolabially and 4) possibly the rare presence of doubled-anterotropid on the lower molars. In contrast,G. duplexandGliruloidesaff.G. duplexnormally have incomplete endoloph on M1/2 and rarely have hook-like protoconid and mesoconid. In both forms, the metalophid usually extends mesiolabially and a doubledanterotropid is common on the lower molars.

    Measurements (length × width in mm) P4 (V 18111.1) 0.87 × 1.05; M2 (V 18110.1) 1.05 × 1.25; M1/2 (V 18112) 1.03 × 1.09; M3 (V 18110.2) 0.92 × 1.05; M3 (V 18113) 0.77 × 0.98; p4 (V 18111.2) 0.86 × 0.68; p4 (V 18111.3) 0.84 × 0.67; m1 (V 18110.3) 1.17 × 1.05; m1 (V 18111.4) 1.16 × 1.09; m1 (V 18110.4) 1.13 × 1.06; m3 (V 18110.5) 1.11 × 1.00 (Except P4 is larger and m3 is slightly longer than inG. duplex, the other teeth are approximate to those ofG. duplexin size.

    Description The occlusal surface of all cheek teeth is concave. All the upper cheek teeth have usually nine transverse ridges, including the six main ridges (anteroloph, protoloph, metaloph, posteroloph, precentroloph and postcentroloph) and three extra ones (anterotrope, prototrope and posterotrope). Both the anterotrope and posterotrope extend almost the full length of the valleys where they locate. The labial ends of the main ridges are free or lightly connected. The lingual wall of the endoloph on upper cheek teeth is well decorated. All upper cheek teeth have three roots (one major lingual and two minor labial ones).

    P4 The left P4 (V 18111.1; Fig. 2A) is suboval-shaped in occlusal view. Its occlusal surface is strongly concave. The anteroloph has its labial end separated from the protoloph and the lingual end touched the protoloph but not the endoloph. The lingual end of the posteroloph is weakly in contact with the endoloph and labially connects to the metaloph. The protoloph is labially free and lingually connects to the endoloph; it is interrupted midway by the U-shaped anterotrope-prototrope connection. The anterotrope extends to the labial border and stays free, whereas the prototrope is half long the anterotrope. The precentroloph ends free labially, but lingually it bifurcates to join the protoloph and metaloph, respectively. The postcentroloph is short and situated in the middle of the valley, with both ends being free. The metaloph is convex distally and interrupted near its lingual end. The posterotrope is long, fi lling nearly thewhole length of the corresponding valley. The trigon is V-shaped.

    Fig. 2Gliruloides zhouigen. et sp. nov. A. P4 sin. IVPP V 18111.1; B. M1/2 sin. V 18112; C. M2 sin. V 18110.1, holotype; D. M3 sin. V 18110.2, paratype; E. M3 dex V 18113; F. p4 dex, V 18111.2; G. p4 dex, V 18111.3; H. m1 dex, V 18110.3, paratype; I. m1 dex, V 18111.4; J. m1 sin., V 18110.4, paratype; K. m3 dex., V 18110.5, paratype

    M1/2 Two M2 and one M1/2 are in the collection. The identi fi cation of specimen V 18112 (Fig. 2B) as M1 or M2 cannot be certain so that we denote it as M1/2 (but is probably M1 because of its V-shaped trigon). The holotype M2 (V 18110.1) is moderately worn and almost identical to the specimen V 11812.1 reported by Wu et al. (2000: pl. 1, fi g. 1). The occlusal surface is wider than long. The paracone and metacone are prominent. The anteroloph is labially weakly connects to the paracone with a shallow furrow in between and lingually joins the endoloph. The paracone connects the labial end of the precentroloph loosely. The metacone is separated anteriorly from the postcentroloph and posteriorly fromthe posteroloph by a narrow and very shallow furrow, respectively. The precentroloph is long, extending to the point near the endoloph. The trigon is narrowly U-shaped and contains the long prototrope, precentroloph and postcentroloph. The lingual wall of the endoloph is well decorated. Specimen V 18112, a left M1/2 is slightly worn and closely similar to the holotype, but differs from the latter in being wider anteriorly than posteriorly, the V-shaped trigon, precentroloph being labially far separated from the paracone and curving backwards to the metacone, the anteroloph being labially free from the paracone, and loosely contacting with the protocone lingually to form a nearly complete endoloph. The anterotrope and posterotrope on all three specimens extend the whole length of the corresponding occupied valley.

    M3 There are three M3 specimens (V 11812.2, V 18110.2 and V 18113), which are characterized by a trapezoid-shape, being much wider anteriorly than posteriorly. M3 has usually the same number of ridges as M1/2 but the precentroloph is much shorter than the postcentroloph. Specimen V 18110.2 is well worn. The labial side of the paracone was slightly damaged but it seems in connection with the anteroloph and protoloph. The precentroloph is much shorter than the postcentroloph and extends labially to the labial margin of the tooth and has no contact with both the paracone and metacone. The postcentroloph extends lingually and almost reaches the endoloph; its labial end is weakly connected with the metacone. The prototrope is absent, instead is a metatrope present inside the trigon. The metatrope is short, only a half width of the valley, and locates labially. Both metaloph and posteroloph are interrupted by a narrow gap at the midway. It seems that the endoloph is separated from the lingual end of the anteroloph before the tooth was worn. The specimen V 11812.2 (Wu et al., 2000: pl. 1, fi g. 2) is highly similar to V 18110.2. There is, in addition to the anterotrope, an extra ridge between the anterotrope and protoloph. Both prototrope and metatrope are present. The specimen V 18113 (Fig. 2E) from XJ 20004 is similar to specimen V 11812.2, but its ridges are more slender and the anterotrope is weaker and shorter. We tentatively assign this tooth to this species.

    The p4 Both p4 are round trapezoid in shape, wider posteriorly than anteriorly (Fig. 2F, G). The anterolophid is convex mesially and connected distally to the metalophid to form a closed loop. The centrolophid is thin and long, connected lingually to the metaconid weakly, and extends to the labial border. The labial end of the mesolophid turns abruptly mesially along the labial border on one specimen (V 18111.2) but becomes well-swollen labially on the other (V 18111.3); the mesolophid connects lingually to the posterolophid at the entoconid, and labially connected to or separated from the posterolophid. The anterotropid and posterotropid are well developed, and the mesotropid is short or long. The metatropid is absent in both teeth so that p4 is eight-ridged.

    The m1 Four specimens are in the collection, including the one described by Wu et al. (2000: pl. 1, fi g. 3). They are quite monotonous morphologically and generally trapezoidshaped with the anterior end being slightly narrower than the posterior end. The m1 usually has nine transverse ridges, including fi ve main ridges (anterolophid, metalophid, centrolophid,mesolophid and posterolophid) and four extra ridges (anterotropid, metatropid, mesotropid and posterotropid). All four extra ridges are well developed, of which the anterotropid and posterotropid extend nearly the entire length of the located valleys. However, specimen V 11812.3, described by Wu et al. in 2000, remains an exception because it has an additional secondary extra ridge between the anterotropid and metalophid. For all specimens of m1, the anterolophid is slightly concave mesially, labial end slightly curved distally and free-ended; its lingual end slightly curved distally too, and gradually merges into the metaconid lingually. The metalophid is lingually connected to the metaconid and extends slightly distolabially, with its labial end turning abruptly mesially to form a hook-like protoconid that is not connected to the anterolophid. The centrolophid is long and extends labially near the mesoconid but does not reach to the labial border of the tooth. Lingually the centrolophid is either separated from or connected with the metaconid. The mesolophid is slightly convex distally and extends mesiolabially where, like the metalophid, it turns abruptly mesially and forms a hook-like mesoconid that is separated from the protoconid mesially and from the hypoconid distally. The posterolophid joins the mesolophid lingually at the entoconid. An endolophid is absent.

    The m3 This tooth is mesially much wider than distally. The anterolophid is slightly convex mesially. Except for the four extra ridges seen in m1, there are two more secondary extra ridges: one mesial and the other distal to the posterotropid, respectively. Furthermore, a very small and weak enamel bulge but not ridge is present between the anterolophid and anterotropid.

    All lower cheek teeth have two roots, but the mesial root of m1 (V 11812.3) shows a trend of bifurcation at the end.

    Several teeth display distinctive striations of abrasion in mesiolingual-distolabial direction on the occlusal surface (Fig. 3), which we record here. We think this information should be useful for further study of this animal in masticatory movement, dietary, and perhaps taxonomy.

    Fig. 3 Occlusal view of the lower and upper cheek teeth ofGliruloides zhouiArrows indicate the direction of striations caused by wear on the concave crown surface

    Comparisons and discussions As mentioned above, the Junggar glirid, represented by three specimens at the early time, was fi rst assigned toGlirulus(Wu et al., 2000) because it is similar toGlirulusin having usually nine ridges on both upper and lower cheek teeth, withwell-developed anterotrope and posterotrope in upper cheek teeth and the anterotropid and posterotropid in lower cheek teeth. However, additional specimens collected subsequently made us to reconsider the taxonomic assignment of the Junggar glirid. We have listed above several features that differ the Junggar form fromGlirulusin the differentiate diagnoses. At the same time, we noted that the Junggar form is quite similar to the Turkish Lower Miocene dormouse,Vasseuromys duplex, in both upper and lower molars. Thus, we considered that the Junggar glirid should not be assigned toGlirulusand have referred it toVasseuromysin several of our papers (Meng et al., 2001, 2006; Ye et al., 2001a, b; 2003a, b) without explaining the reason until this study.

    Vasseuromyswas established by Baudelot and de Bonis (1966) based on the type speciesV. rugosusfrom the Lower Miocene of Laugnac, France. The only material of this species was a mandible with p4-m2. The original generic diagnosis states (translated from French by Daams and de Bruijn, 1995:50): “Medium-sized Gliridae. Cheek teeth with concave occlusal surface. Lower molars characterized by a centrolophid reaching the labial border and by a longitudinal prolongation of the labial cusps that form a nearly continuous ectolophid”. Based on additional specimens of upper cheek teeth from the type locality de Bonis (1973:54) emended the diagnosis of the genus as: “Vasseuromyscharacterized by multiplicity of the extra ridges and by the upper molars with a continuous endoloph (translated from French)”. The upper cheek teeth referred in the work of de Bonis (1973) consisted of one P4, one M1/2, and two M3. While working on the dormice from Austria and Spain, Daxner-H?ck and de Bruijn (1981) and Alvarez Sierra et al. (1990) have independently visited the collections ofVasseuromys rugosushoused at the University of Utrecht; all the specimens are from the type locality. The two research teams noticed that there were several upper molars ofV. rugosuswith incomplete endoloph. Alvarez Sierra et al. (1990) wrote: “Although de Bonis (1973) characterizes this species as having a continuous endoloph (on the basis of one worn specimen), several M1,2 without endoloph are present in the collections of the University of Utrecht. The specimens from the Utrecht collections have a long anteroloph whose lingual end descends toward the base of the protocone.” Daxner-H?ck and de Bruijn (1981) further provided fi gures of the upper and lower molars ofV. rugosus, which show the V-shaped or narrow U-shaped trigon and incomplete endoloph in the upper molars (Daxner-H?ck and de Bruijn, 1981: fi g. 1-h,r,s,t,u; fi g. 2-m,n,o,p) (see Fig. 4B).

    Up to now ten species have been included in this genus (ref. Daams and de Bruijn, 1995; Ruiz-Sánchez et al., 2012a, b; 2014):

    Vasseuromys rugosusBaudelot & de Bonis, 1966; type locality and type level: Laugnac of France, MN2B, Early Miocene.

    V. priscusde Bonis, 1973; type locality and type level: Moissac 1 of France, MN1, Early Miocene.

    V. pannonicus(Kretzoi, 1978); type locality and type level: Budapest, Freshwater Limestone of the Széchenyi hill; MN10?, Late Miocene (Synonym:V. theniiDaxner-H?ck &de Bruijn, 1981).

    Fig. 4 The comparison of the upper and lower cheek teeth ofGliruloides zhoui,Vasseuromys rugosus,Glis guerbuezi, andGlirulus japonicusA.Gliruloides zhouigen. et sp. nov. from northern Junggar Basin, Late Oligocene (M2 and m2); B.Vasseuromys rugosusBaudelot & de Bonis, 1966 from the type locality Laugnac of France, MN2B, Early Miocene (after Daxner-H?ck and de Bruijn, 1981) (P4-M3 and p4-m3); C.Glis guerbueziünay-Bayraktar, 1989 from Thrace of Turkey (modi fi ed from ünay-Bayraktar, 1989) (M2 and m2); D.Glirulus japonicus(Schinz, 1845) from the upper part of Horizon M of Kannondo Cave Site of Japan (modi fi ed from Kawamura, 1989) (M1 and m2)

    V. autolensis(Cuenca, 1985); type locality and type level: Autol, La Rioja, Spain; MN1, Early Miocene.

    V. bacchius(Martínez-Salanova, 1987); type locality and type level: Fuenmayor 2, Autol 1, La Rioja, Spain; MN2B, Early Miocene.

    V. elegansWu, 1993; type locality and type level: Stubersheim 3, Germany, MN3, Early Miocene.

    V. duplexünay, 1994; type locality and type level: Harami 1, Turkey; MN2, Early Miocene.

    V. cristinaeRuiz-Sánchez et al., 2012; type locality and type level: Pico del Fraile 2, Ebro Basin, Spain; MN4/5, Miocene.

    V. rambliensisRuiz-Sánchez et al., 2012; type locality and type level: Pico del Fraile 1, Ebro Basin, Spain; Upper Ramblian, MN3, zone A, Early Miocene

    V. bergasensisRuiz-Sánchez et al., 2014; type locality and type level: Bergasa, Ebro Basin, Spain; MP30, Late Oligocene (ref. Lacomba, 1988).

    Apart from the above listed species,Nievella mayriDaams, 1976 from the Early Miocene of Cetina de Aragón, the early Late MioceneRamys multicresatus(de Bruijn, 1966) andMyolidusmay belong toVasseuromys, as so suggested by Agusti et al. (2011).

    The taxonomy, phylogeny and biostratigraphy of the known species ofVasseuromysare beyond the scope of this study. Our focus is on whether the Junggar form belongs toVasseuromys, a potential assignment we have indicated in our previous studies (Meng et al., 2001, 2006; Ye et al., 2001a, b; 2003a, b).

    After intensive survey of the literatures aboutVasseuromys, we realized that, in general, all European species have features in common with the type speciesV. rugosusand differ from those of the TurkishV. duplexandVasseuromysaff.V. duplex. We recognize the fundamental difference between them as representing two dental patterns ofVasseuromyscheek teeth, primarily re fl ected in the upper cheek teeth, which are differentiated below:

    In European species ofVasseuromys, the anterotrope and posterotrope outside the trigon are usually absent; if present, they are short and weak; the ridges are rugose and asymmetrically arranged.

    InV. duplexandVasseuromysaff.V. duplexfrom Anatolia of Turkey, the anterotrope and posterotrope outside the trigon are always present and long; they extend almost the full length of the corresponding valleys they are in; the ridges are regular and symmetrically arranged.

    In fact, ünay (1994:470) already noted that “The striking features of TurkishVasseuromysare the ever present long extra ridges in the anterior and posterior valleys outside the trigon on the M1/2… different from all European species.” We found that the pattern of the anterotrope and posterotrope is present not only on M1/2 but also on P4 and M3 of TurkishVasseuromys(ünay, 1994). Because of the distinct morphological differences between the Turkish and EuropeanVasseuromysspecies, we think thatV. duplexshould be inevitably assigned to a different genus. The Junggar and Turkish forms are also different fromGlirulus, as we discussed above.

    In addition to the tooth crown structures, we have compared the root number of various species ofGlirulus(Paraglirulusincluded),GliruloidesandVasseuromys(Table 1). All upper cheek teeth of the three genera are triple-rooted except thatGlirulus(Paraglirulus)werenfelsihas a double-rooted P4. However, the available data show that the lower cheek teeth of allGlirulusspecies are double-rooted except that p4 ofGlirulus(P.)agelakifrom Aliveri issingle-rooted andGlirulus japonicushas double-rooted p4 and triple-rooted lower molars.Gliruloideshas also double-rooted lower cheek teeth except that the m3 ofGliruloidesaff.G. duplexis occasionally triple-rooted. Species ofVasseuromys, where the root condition is known, however, have a single-rooted p4 and double or triple-rooted lower molars. Table 1 shows that the Junggar form is similar toG. duplexnot only in their tooth crown structures but also in the root number. However, it should be noted that only the m3 root number (two or three roots) ofGliruloidesaff.G. duplexwas provided by ünay (1994), whereas the root number of m1/2 was not provided. With the survey of the root number distributions among relevant species, we consider the root condition to be of taxonomic information for species of the three genera in question, although more data are needed to verify this consideration.

    In short, given that the Junggar and Turkish forms share some distinctive dental features but differ from theVasseuromysandGlirulusin various features, we think they should beplaced in the same genus; thus we establish the new genusGliruloidesto include the Junggar and Turkish forms.

    Table 1 Comparisons of cheek teeth root number ofGlirulus,GliruloidesandVasseuromys

    Gliruloidescurrently includes three species:G. zhoui,G. duplexandGliruloidesaff.G. duplexand tentatively assigned to the subfamily Dryomyinae because of several diagnostic features, such as the concaved surface, basically symmetrical upper molar, and some teeth with nearly complete endoloph.

    The Origin ofGliruloidesThe Early Oligocene (ca. MP25)Glis guerbuezifrom Thrace of Turkey (ünay-Bayraktar, 1989) is, to our knowledge, the earliest known dormouse that displays considerable resemblance toGliruloidesin dental pattern.G. guerbuezihas at most nine transverse ridges in either upper or lower molars (see Fig. 4C). On certain M1/2s ofG. guerbuezi, besides four main ridges, there are precentroloph, postcentroloph and prototrope inside the trigon, and anterotrope and posterotrope outside the trigon. In addition to four main ridges and centrolophid, the anterotropid and posterotropid are developed in m1 and m2, with incipient metatropid and mesotropid present in m2. M1/2 ofG. guerbuezialso has V-shaped or narrow U-shaped trigon and an incomplete endoloph, and the anterolophid is labially isolated in lower molars. It is probable thatGliruloideswas derived from aGlis guerbuezilike ancestor, although we are not able to illustrate their detailed evolutionary process because of the limited knowledge available to us. Together withGliruloidesduplexandGliruloidesaff.G. duplex, ünay (1994) also described two species ofGlirulus:G. ekremiof MN3 (from Kesek?y) andGlirulusaff.G. ekremiof MN2 (from Harami 1, associated withV. duplex)) from Anatolia. We have noticed that the TurkishG. ekremihas an incomplete endoloph in 27.50% of M1/2 specimens and a precentroloph connected to the endoloph in only 19.2% of M1/2s, and the lower molars ofG. ekremiare morphologically very similar to those of the Turkish “Vasseuromys”. Thus, identi fi cation of those teeth is not always unquestionable. It is also reasonable to infer thatGlirulusmay also be derived from the sameGlis guerbuezi-like ancestor so thatGliruloidesandGlirulusshare a most recent common ancestor but evolved in different directions as two lineages. The fact that the root number of the mostGlirulusspecies is same as that ofGliruloides, as shown in Table 1, could be another evidence for their common origin.

    Ecology According to Walker (1975: volume 2:979) the livingGlirulus japonicusinhabits mountain forests from about 400 to 1800 meters in elevation, with the highest recorded elevation being 2900 meters. This animal’s diet includes fruits, seeds, insects and bird’s eggs. All fossil specimens ofGliruloidesduplex,Gliruloidesaff.G. duplexand two species ofGlirulusünay (1994) described were from Lower Miocene lignite containing sections (de Bruijn and Sara?, 1991) of Anatolia (from localities Kesek?y, Harami 1-3 and Kil?ak 0, 0", 3A, 3B). The morphological similarity of teeth inGliruloidesandGlirulusfrom Turkey could be attributed to a similar ecological environment (ecotope) and diet. The presence of lignite in the sections probably indicates wet biotopes (de Bruijn and Sara?, 1991; ünay, 1994).Gliruloides zhouiis a member of the Late Oligocene Tieersihabahe-Izone fauna, collected from the same localities (XJ 20004 and XJ 98024) as the giant rhinoParaceratherium sui(Ye et al., 2003a) but was discovered from the level immediately above the level where rhinoceros located. Three rhino localities were found in Ulungur River area, which arrange in a west-east extending line with the largest distance of ca. 57 kilometers.Paraceratherium suiwas discovered from the fl uvial sediments at the base of Tieersihabahe Formation. Ye et al. (2012:1530) inferred that the presence of this giant rhino, at the time of ca. 25 Ma, is an implication of a temperature rise in the course of global cooling and aridi fi cation that begins from the Early Oligocene, which is coincided with the time of “Late Oligocene warming” (Zachos et al., 2001: fig. 2).Gliruloides zhouimight still lived in a relatively wet and warm transitional time somewhat later thanParaceratherium sui. We have mentioned above thatGliruloidescould be derived from theGlis guerbuezi-like form. The latter came from the Turkish late Early Oligocene localities situated in the Lignite-Sandstone Formation (ünay-Bayraktar, 1989:10). The sediments could also indicate that wet and forest environment existed during that period in this regional area.

    Acknowledgements We thank Dr. Ruiz-Sánchez from Department of Geology, Division of Paleontology, University of Valencia of Spain for providing us important literatures about spanishVasseuromys. Thanks are also due to the American Museum of Natural History, New York for permission to use the scanning electron microscope. We are grateful to Su Jianfen and Wu Shaoyuan for their hard work in the fi eld and to Yue Qiwan for sorting the miniscule teeth from the concentration of screenwashed samples. We also thank many local peoples who helped us in collecting, transporting and screenwashing dirt samples under the baking sun of the desert. This research has been supported by the Strategic Priority Research Program of Chinese Academy of Sciences (CAS, XDB03020501) and National Basic Research Program of China (2012CB821904).

    Agustí J, Pérez-Rivarés F J, Cabrera L et al., 2011. The Ramblian-Aragonian boundry and its signi fi cance for the European Neogene continental chronology. Contributions from the Ebro Basin record (NE Spain). Geobios, 44(2-3): 121-134

    Alvarez-Sierra M A, Daams R, Lacomba J I et al., 1990. Paleontology and biostratigraphy (micromammals) of the continental Oligocene-Miocene deposits of the North-Central Ebro Basin (Huesca, Spain). Scripta Geol, 94: 1-77

    Baudelot S, Bonis L de, 1966. Nouveaux Gliridés (Rodentia) de l’Aquitanien du basin d’Aquitaine. C R Soc Géol France, 8: 303-304

    Bonis L de, 1973. Contribution a l’etude des mammifères de l’Aquitanien de l’Agenais: Rongeurs-Carnivores-Perissodactyles. Mém Mus Natl Hist Nat, Ser C, 28: 1-192

    Bruijn H de, 1966. On the mammalian fauna of theHipparion-beds in the Calatayud-Teruel Basin (Prov. Zaragoza, Spain). The Gliridae. Proc K Ned Akad Wet, Ser B, 69(3): 367-387

    Bruijn H de, Sara? G, 1991. Early Miocene rodent faunas from the eastern Mediterranean area, Part I. The genusEumyarion. Proc K Ned Akad Wet, Ser B, 94(1): 1-36

    Cuenca G, 1985. Los Roedores (Mammalia) del Mioceno Inferior de Autol (La Rioja). Cienc Tierra, 2: 1-96

    Daams R, 1976. Miocene rodents (Mammalia) from Cetína de Aragon (Prov. Zaragoza) and Bu?ol (Prov. Valencia), Spain. Proc K Ned Akad Wet, Ser B, 79(3): 152-182

    Daams R, Bruijn H de, 1995. A classi fi cation of the Gliridae (Rodentia) on the basis of dental morphology. Histryx, 6(1-2): 3-50

    Daxner-H?ck G, Bruijn H de, 1981. Gliridae (Rodentia, Mammalia) des Eichkogels bei M?dleng (Nieder?sterreich). Pal?ont Z, 55(2): 157-172

    Daxner-H?ck G, H?ck E, 2009. New data on Eomyidae and Gliridae (Rodentia, Mammalia) from Late Miocene of Austria. Ann Naturhist Mus Wien, 111A: 375-444

    Freudenthal M, 2004. Gliridae (Rodentia, Mammalia) from the Eocene and Oligocene of the Sierra Palomera (Teruel, Spain). Treb Mus Geol Barcelona, 12: 97-173

    Kawamura Y, 1989. Quaternary rodent faunas in the Japanese Islands (part 2). Mem Fac Sci, Kyoto Univ, Ser Geol Mineral, 54(1-2): 125-157

    Kretzoi M, 1978. Wichtigere Streufunde in der Wirbeltierpal?ontologischen Sammlung der Ungarischen Geologischen Anstalt. áll F?ldt Int évi Jel, 1978: 348-358

    Lacomba J I, 1988. Estudio de las faunas de micromamiferos del Oligoceno superior y mioceno inferior en las cuencas deLoranca, Ebro riojano y Ebro aragonés. Aspectos paleoecológicos. Doctoral thesis. Madrid: Madrid Universidad Complu tense de Madrid. 1-389

    Martínez-Salanova J, 1987. Estudio paleontológico de los Micromamiferos del Mioceno inferior de Fuenmayor (La Rioja). Cienc Tierra, 10: 1-99

    Mayr H, 1979. Gebissmorphologische Urtersuchungen an miozaenen Gliridae (Mammalia, Rodentia) Süddeutshlands. Inaugural Dissertation. München: Ludwig-Maximilians-Universit?t. 1-380

    Meng J, Ye J, Wu W Y et al., 2001. Two petrosals of gliriform mammals from Late Oligocene of Tieersihabahe, Xinjiang Uygur Autonomous Region, China. Vert PalAsiat, 39(1): 43-53

    Meng J, Ye J, Wu W Y et al., 2006. A recommended boundary stratotype section for Xiejia Stage from northern Junggar Basin: implications to related bio-chronostratigraphy and environmental changes. Vert PalAsiat, 44(3): 205-236

    Meulen A J van der, Bruijn H de, 1982. The mammal from the Lower Miocene of Aliveri (Island of Evia, Greece). Part 2, The Gliridae. Proc K Ned Akad Wet, Ser B, 85(4): 485-524

    Ruiz-Sánchez F J, Murelaga X, Freudenthal M et al., 2012a. A new species of glirid rodentVasseuromys(Gliridae, Rodentia) from the Aragonian (Miocene) of the Ebro Basin (north-eastern Spain). Acta Palaeont Pol, 57(2): 225-239

    Ruiz-Sánchez F J, Murelaga X, Freudenthal M et al., 2012b.Vasseuromysrambliensissp. nov. (Gliridae, Mammalia) from the Ramblian (Lower Miocene) of the Tudela Formation (Ebro Basin, Spain). Paleont Electron, 15(1), 4A: 1-16

    Ruiz-Sánchez F J, Lacomba-Andueza J I, Freudenthal M et al., 2014. A new species ofVasseuromys(Gliridae, Mammalia) from the Upper Oligocene of the Ebro Basin (Spain). Pal?ont Z, 88(1): 73-84

    ünay E, 1994. Early Miocene rodent faunas from the eastern Mediterranean area. Part IV. The Gliridae. Proc K Ned Akad Wet, Ser B, 97(4): 445-490

    ünay-Bayraktar E, 1989. Rodents from the Middle Oligocene of Turkish Thrace. Utrecht Micropal Bull Spec Publ, 5: 1-119

    Walker E P, 1975. Mammals of the World. 3rd ed. Baltimore and London: The Johns Hopkins University Press. 1-1500

    Wu W Y, 1993. Neue Gliridae (Rodentia, Mammalia) aus untermiozanen (orleanischen) Spaltenfüllungen Süddeutschlands. Doc Nat, 81: 1-157

    Wu W Y, Ye J, Bi S D et al., 2000. The discovery of Late Oligocene dormice from China. Vert PalAsiat, 38(1): 36-42

    Ye J, Meng J, Wu W Y, 2003a. Discovery ofParaceratheriumin the northern Junggar Basin of Xinjiang. Vert PalAsiat, 41(3): 220-229

    Ye J, Meng J, Wu W Y, 2003b. Oligocene/Miocene beds and faunas from Tieersihabahe in the northern Junggar Basin of Xinjiang. Bull Am Mus Nat Hist, 279: 568-585

    Ye J, Wu W Y, Meng J, 2001a. Tertiary stratigraphy in the Ulungur River Area of the northern Junggar Basin of Xinjiang. J Stratigr, 25(3): 193-200

    Ye J, Wu W Y, Meng J, 2001b. The age of Tertiary strata and mammal faunas in Ulungur River Area of Xinjiang. J Stratigr, 25(4): 283-287

    Ye J, Wu W Y, Ni X J et al. 2012. The Duolebulejin section of northern Junggar Basin and its stratigraphic and environmental implication. Sci Sin Terrae, 42: 1523-1532

    Zachos J, Pagani M, Sloan L et al., 2001. Trends, rhythms and aberrations in global climate 65 Ma to Present. Science, 292: 686-693

    新疆準噶爾盆地北緣晚漸新世睡鼠再研究

    吳文裕1孟 津1,2葉 捷1倪喜軍1畢順東1,3

    (1 中國科學院古脊椎動物與古人類研究所,中國科學院脊椎動物演化與人類起源重點實驗室 北京 100044)
    (2 美國自然歷史博物館古生物學部 紐約 10024)
    (3 美國賓夕法尼亞州印第安納大學生物系 印第安納 PA 15705)

    周氏似日本睡鼠Gliruloides zhoui是發(fā)現(xiàn)于新疆準噶爾盆地北緣晚漸新世鐵爾斯哈巴合哺乳動物組合I帶的一個化石新屬種。新屬似日本睡鼠Gliruloides的屬征為:中等大小;頰齒咀嚼面凹;上、下頰齒通常具9條主要橫脊,有時具次級附脊;上頰齒的前邊附脊和后邊附脊以及下頰齒的下前邊附脊和下后邊附脊都很發(fā)育,幾乎占據(jù)了其所在齒谷的整個長度。上頰齒的橫脊唇端趨于游離。M1和M2具V形或窄U形三角座,內脊不完整或近于完整,前中央脊不與內脊相連。下頰齒的下內脊通常不連續(xù);下臼齒的下前邊脊在唇端稍向后彎,但不與原尖相連。p4, m1-m3, P4, M1-M3的齒根數(shù)分別為2, 2, 3, 3。新屬與Glirulus在形態(tài)上相似,但Glirulus的上頰齒的三角座均為寬U型,具有完整的內脊,前中央脊通常與內脊相交,橫脊唇端通常不游離。新屬與Vasseuromys屬的最主要形態(tài)差異在于上頰齒具有很發(fā)育的前邊附脊和后邊附脊,而后者上頰齒的前邊附脊和后邊附脊通常缺失或很不發(fā)育。歸入該屬的種還有土耳其早中新世的Vasseuromysduplex和Vasseuromysaff.V. duplex。土耳其Thrace早漸新世的Glis guerbuezi很可能是Gliruloides和Glirulus的共同祖先類型。Gliruloides可能生活于溫濕的生態(tài)環(huán)境。

    準噶爾盆地北緣,晚漸新世,鐵爾斯哈巴合哺乳動物組合I帶,睡鼠

    Q915.873

    A

    1000-3118(2016)01-0036-15

    2015-07-22

    Wu W Y, Meng J, Ye J et al., 2016. Restudy of the Late Oligocene dormice from northern Junggar Basin. Vertebrata PalAsiatica, 54(1): 36-50

    中國科學院戰(zhàn)略性先導科技專項(編號:XDB03020501)和國家重點基礎研究發(fā)展計劃項目(編號:2012CB821904)資助。

    猜你喜歡
    印第安納哈巴準噶爾盆地
    靳叔家的哈巴
    百花園(2022年6期)2022-12-28 04:49:34
    印第安納大學玻璃建筑——跨越70年的密斯·凡·德·羅“新作”
    房地產導刊(2022年5期)2022-06-01 06:19:24
    15 個產地玄參中哈巴苷與哈巴俄苷含量測定
    準噶爾盆地八道灣組濕地扇三角洲沉積特征
    哈巴雪山的太陽
    民族音樂(2018年2期)2018-05-26 03:04:34
    準噶爾盆地南緣齊古背斜復雜構造模式研究
    新疆地質(2016年4期)2016-02-28 19:18:43
    準噶爾盆地南緣泥火山與油氣苗
    石油知識(2016年2期)2016-02-28 16:19:48
    準噶爾盆地西北緣克-夏斷裂帶構造特征新認識
    新疆地質(2015年3期)2015-12-10 05:08:27
    微波輔助萃取法提取玄參中哈巴苷和哈巴俄苷的工藝研究
    高校應急管理機制的構成及運作——以美國印第安納大學為例
    亚洲,欧美,日韩| 午夜av观看不卡| 爱豆传媒免费全集在线观看| 在线观看免费高清a一片| 亚洲精品成人av观看孕妇| 亚洲国产看品久久| 黄色一级大片看看| 国产精品一国产av| 制服人妻中文乱码| 51国产日韩欧美| 久久精品国产a三级三级三级| 日韩在线高清观看一区二区三区| 黑丝袜美女国产一区| 中文字幕人妻丝袜制服| 狠狠婷婷综合久久久久久88av| 日韩,欧美,国产一区二区三区| 欧美日本中文国产一区发布| 亚洲经典国产精华液单| 免费观看av网站的网址| 亚洲av中文av极速乱| 婷婷色综合大香蕉| 十分钟在线观看高清视频www| 日韩av免费高清视频| 亚洲av电影在线观看一区二区三区| 久久青草综合色| 9191精品国产免费久久| 国产精品嫩草影院av在线观看| 欧美 日韩 精品 国产| 人人妻人人爽人人添夜夜欢视频| av播播在线观看一区| 看非洲黑人一级黄片| 久久人人爽人人爽人人片va| 中文乱码字字幕精品一区二区三区| 亚洲少妇的诱惑av| 深夜精品福利| 国产福利在线免费观看视频| 桃花免费在线播放| 激情五月婷婷亚洲| 狂野欧美激情性bbbbbb| 一二三四在线观看免费中文在 | 午夜精品国产一区二区电影| 飞空精品影院首页| 丝袜在线中文字幕| 91成人精品电影| 亚洲人成网站在线观看播放| 久久精品人人爽人人爽视色| 男的添女的下面高潮视频| 婷婷色麻豆天堂久久| 国产精品国产三级专区第一集| 婷婷色综合大香蕉| 一本—道久久a久久精品蜜桃钙片| 精品久久国产蜜桃| 亚洲精品久久午夜乱码| 性色avwww在线观看| 国产片特级美女逼逼视频| 丝袜脚勾引网站| 欧美亚洲日本最大视频资源| 丝袜喷水一区| 亚洲国产精品999| 色94色欧美一区二区| 精品人妻偷拍中文字幕| 成人18禁高潮啪啪吃奶动态图| 美女国产高潮福利片在线看| 18禁国产床啪视频网站| 最黄视频免费看| 一区二区三区四区激情视频| 新久久久久国产一级毛片| 人体艺术视频欧美日本| 亚洲av男天堂| 国产精品久久久久成人av| 精品人妻一区二区三区麻豆| 寂寞人妻少妇视频99o| xxx大片免费视频| 2021少妇久久久久久久久久久| 男女午夜视频在线观看 | 9热在线视频观看99| 日韩电影二区| 2022亚洲国产成人精品| 一区二区三区四区激情视频| 日日啪夜夜爽| 22中文网久久字幕| 中国国产av一级| a级毛色黄片| 在线观看免费视频网站a站| 久久久久久久久久久久大奶| 黑人猛操日本美女一级片| 狂野欧美激情性bbbbbb| 亚洲精品美女久久av网站| 免费在线观看黄色视频的| 丝袜美足系列| 日韩视频在线欧美| 日韩av在线免费看完整版不卡| 免费av中文字幕在线| 水蜜桃什么品种好| 欧美激情国产日韩精品一区| 少妇人妻久久综合中文| 丰满少妇做爰视频| 91成人精品电影| 午夜福利在线观看免费完整高清在| av又黄又爽大尺度在线免费看| 亚洲国产色片| 人妻一区二区av| 男人操女人黄网站| 亚洲国产最新在线播放| 亚洲第一av免费看| 婷婷色麻豆天堂久久| 天堂8中文在线网| 亚洲国产成人一精品久久久| 九九在线视频观看精品| 免费av中文字幕在线| 韩国av在线不卡| 亚洲成av片中文字幕在线观看 | 国产欧美另类精品又又久久亚洲欧美| 狂野欧美激情性bbbbbb| 国产欧美亚洲国产| 各种免费的搞黄视频| 亚洲国产精品成人久久小说| 人人妻人人爽人人添夜夜欢视频| 久久这里只有精品19| 性色avwww在线观看| 国产又爽黄色视频| 国产精品偷伦视频观看了| 日本爱情动作片www.在线观看| 久久久久久久国产电影| 日本黄大片高清| 夫妻午夜视频| 久久久精品区二区三区| 国产男女超爽视频在线观看| 有码 亚洲区| 久久人人爽人人片av| 十八禁网站网址无遮挡| av又黄又爽大尺度在线免费看| 最近2019中文字幕mv第一页| 免费大片黄手机在线观看| 少妇人妻久久综合中文| 九色亚洲精品在线播放| 巨乳人妻的诱惑在线观看| 成年动漫av网址| 最近手机中文字幕大全| 色婷婷久久久亚洲欧美| 国产精品一二三区在线看| 伊人久久国产一区二区| 亚洲,欧美,日韩| 国产永久视频网站| 最近最新中文字幕免费大全7| 久久国产亚洲av麻豆专区| 久久久久国产精品人妻一区二区| 丝袜喷水一区| 亚洲人成77777在线视频| 少妇被粗大猛烈的视频| 国产视频首页在线观看| 曰老女人黄片| 丁香六月天网| 国产精品一区二区在线观看99| av免费观看日本| 国产日韩一区二区三区精品不卡| 久久久久精品性色| 国产在线一区二区三区精| 九草在线视频观看| 日本wwww免费看| 国产免费一区二区三区四区乱码| 日韩在线高清观看一区二区三区| 成人国语在线视频| 黄网站色视频无遮挡免费观看| 亚洲中文av在线| 精品久久蜜臀av无| 啦啦啦啦在线视频资源| 香蕉丝袜av| 亚洲婷婷狠狠爱综合网| 在线观看免费视频网站a站| 国产黄频视频在线观看| videossex国产| 久久婷婷青草| 看非洲黑人一级黄片| 在现免费观看毛片| 岛国毛片在线播放| 色哟哟·www| 啦啦啦啦在线视频资源| 欧美xxⅹ黑人| 亚洲国产精品一区三区| 国产精品麻豆人妻色哟哟久久| 久久久亚洲精品成人影院| 久久99热这里只频精品6学生| 国产日韩欧美亚洲二区| 免费人妻精品一区二区三区视频| 丝袜美足系列| 久久久国产欧美日韩av| 欧美激情 高清一区二区三区| 国产无遮挡羞羞视频在线观看| 国产欧美日韩一区二区三区在线| av视频免费观看在线观看| 在线看a的网站| 少妇的丰满在线观看| 免费观看性生交大片5| 欧美xxxx性猛交bbbb| 亚洲欧美色中文字幕在线| 国产av精品麻豆| 少妇人妻 视频| 免费观看无遮挡的男女| 亚洲av综合色区一区| 一本一本久久a久久精品综合妖精 国产伦在线观看视频一区 | 黑人高潮一二区| av电影中文网址| videos熟女内射| 亚洲成色77777| 精品国产一区二区久久| 91在线精品国自产拍蜜月| 国产精品久久久久久久久免| 日韩,欧美,国产一区二区三区| 国产欧美日韩综合在线一区二区| 国精品久久久久久国模美| 熟妇人妻不卡中文字幕| 久久女婷五月综合色啪小说| 你懂的网址亚洲精品在线观看| 欧美成人午夜免费资源| 国产在线视频一区二区| 蜜臀久久99精品久久宅男| 久久人人爽av亚洲精品天堂| 久久久久久久国产电影| 在线天堂中文资源库| 美女主播在线视频| av不卡在线播放| h视频一区二区三区| 日韩人妻精品一区2区三区| 一边亲一边摸免费视频| 亚洲色图综合在线观看| 在线 av 中文字幕| 三级国产精品片| 亚洲国产精品999| 超色免费av| 桃花免费在线播放| 18禁在线无遮挡免费观看视频| 99久国产av精品国产电影| 成人二区视频| 在线精品无人区一区二区三| 欧美精品一区二区免费开放| 搡老乐熟女国产| 水蜜桃什么品种好| 欧美 日韩 精品 国产| 久久鲁丝午夜福利片| 亚洲激情五月婷婷啪啪| 69精品国产乱码久久久| 激情五月婷婷亚洲| av女优亚洲男人天堂| 免费少妇av软件| 久久久国产精品麻豆| 午夜日本视频在线| 女人精品久久久久毛片| av国产久精品久网站免费入址| 熟妇人妻不卡中文字幕| 亚洲成人av在线免费| 国产一区二区在线观看日韩| 欧美日韩视频高清一区二区三区二| 少妇熟女欧美另类| 男女午夜视频在线观看 | 一级毛片电影观看| 国产片特级美女逼逼视频| 丝袜在线中文字幕| 久久狼人影院| 免费大片黄手机在线观看| 欧美亚洲 丝袜 人妻 在线| 黄色怎么调成土黄色| 女的被弄到高潮叫床怎么办| 日韩在线高清观看一区二区三区| 欧美人与善性xxx| 午夜av观看不卡| 美女xxoo啪啪120秒动态图| 97人妻天天添夜夜摸| 久久精品熟女亚洲av麻豆精品| 亚洲,欧美,日韩| 香蕉国产在线看| 18禁在线无遮挡免费观看视频| 天天影视国产精品| 这个男人来自地球电影免费观看 | 国产黄色免费在线视频| 亚洲色图综合在线观看| 丁香六月天网| 中文欧美无线码| 久久精品国产综合久久久 | 18在线观看网站| 亚洲精品,欧美精品| 街头女战士在线观看网站| 中文欧美无线码| 国产精品国产三级国产av玫瑰| 国产高清国产精品国产三级| 美国免费a级毛片| 精品一区二区三区视频在线| 超碰97精品在线观看| 欧美成人午夜免费资源| 搡女人真爽免费视频火全软件| 大片电影免费在线观看免费| 国产亚洲欧美精品永久| 亚洲精品国产色婷婷电影| 国产成人免费观看mmmm| 波多野结衣一区麻豆| 一区二区三区精品91| 黄片播放在线免费| 日韩欧美精品免费久久| 亚洲欧美一区二区三区国产| 日日啪夜夜爽| 一区二区日韩欧美中文字幕 | av网站免费在线观看视频| 精品一区在线观看国产| www日本在线高清视频| 国产xxxxx性猛交| 国产一区二区三区av在线| 国产欧美另类精品又又久久亚洲欧美| 亚洲av电影在线观看一区二区三区| av卡一久久| 亚洲精品国产av蜜桃| 天堂中文最新版在线下载| 69精品国产乱码久久久| av.在线天堂| 亚洲内射少妇av| 欧美丝袜亚洲另类| 一区二区日韩欧美中文字幕 | 国产日韩欧美视频二区| 熟妇人妻不卡中文字幕| 免费观看在线日韩| 在线观看国产h片| 亚洲精品中文字幕在线视频| 精品少妇黑人巨大在线播放| 搡女人真爽免费视频火全软件| 97人妻天天添夜夜摸| 亚洲,欧美精品.| 亚洲美女黄色视频免费看| 啦啦啦中文免费视频观看日本| 丁香六月天网| 男的添女的下面高潮视频| 国产精品久久久av美女十八| 另类亚洲欧美激情| av在线观看视频网站免费| 视频中文字幕在线观看| 蜜桃国产av成人99| 久久久久国产网址| 18禁裸乳无遮挡动漫免费视频| 亚洲精品成人av观看孕妇| 国产精品一区二区在线观看99| 国产激情久久老熟女| 中文字幕亚洲精品专区| 国产免费福利视频在线观看| 天美传媒精品一区二区| 免费黄频网站在线观看国产| freevideosex欧美| 欧美日韩国产mv在线观看视频| 亚洲国产看品久久| 观看av在线不卡| 老司机影院成人| 国产欧美亚洲国产| 国产精品免费大片| 久久人人97超碰香蕉20202| 欧美日韩一区二区视频在线观看视频在线| av天堂久久9| 免费黄频网站在线观看国产| 大香蕉久久成人网| 日韩免费高清中文字幕av| 尾随美女入室| 18禁裸乳无遮挡动漫免费视频| 激情视频va一区二区三区| 中文字幕另类日韩欧美亚洲嫩草| 精品久久国产蜜桃| 亚洲图色成人| 男女边摸边吃奶| 咕卡用的链子| 深夜精品福利| 一级片'在线观看视频| 久久精品夜色国产| 国产69精品久久久久777片| 香蕉精品网在线| freevideosex欧美| 免费日韩欧美在线观看| 国产又色又爽无遮挡免| 国产男女内射视频| 久久久久久人妻| 国产极品天堂在线| 高清欧美精品videossex| 免费看光身美女| 国国产精品蜜臀av免费| www日本在线高清视频| 91精品三级在线观看| 一本—道久久a久久精品蜜桃钙片| 国产精品国产三级国产专区5o| 亚洲伊人色综图| 国产黄色免费在线视频| 99久久综合免费| 国产精品人妻久久久久久| 日日撸夜夜添| 女人被躁到高潮嗷嗷叫费观| 九草在线视频观看| 国产精品人妻久久久影院| 国产一级毛片在线| 天天操日日干夜夜撸| 97在线视频观看| 久久精品国产a三级三级三级| 1024视频免费在线观看| 最后的刺客免费高清国语| 免费在线观看完整版高清| av又黄又爽大尺度在线免费看| 女人被躁到高潮嗷嗷叫费观| 久久久亚洲精品成人影院| 一区二区三区乱码不卡18| 久久久久久久久久成人| 精品午夜福利在线看| 久久精品国产亚洲av天美| 久久99蜜桃精品久久| 美女国产高潮福利片在线看| 波野结衣二区三区在线| 成人18禁高潮啪啪吃奶动态图| 免费少妇av软件| www.色视频.com| 亚洲精品456在线播放app| 视频在线观看一区二区三区| 国内精品宾馆在线| 欧美日韩综合久久久久久| 欧美性感艳星| 成人午夜精彩视频在线观看| 国产极品粉嫩免费观看在线| 一级毛片我不卡| 日本爱情动作片www.在线观看| 国产欧美日韩一区二区三区在线| 精品国产露脸久久av麻豆| 久久人人爽人人爽人人片va| 国产无遮挡羞羞视频在线观看| 亚洲精品aⅴ在线观看| av卡一久久| 国产精品国产av在线观看| 国产成人91sexporn| 妹子高潮喷水视频| 精品福利永久在线观看| 日韩成人av中文字幕在线观看| 中文精品一卡2卡3卡4更新| 蜜桃国产av成人99| 美女大奶头黄色视频| 亚洲精品国产av成人精品| 免费大片黄手机在线观看| 免费观看性生交大片5| 亚洲欧美成人精品一区二区| 久久人人爽人人爽人人片va| 伊人久久国产一区二区| 中文字幕人妻熟女乱码| 欧美3d第一页| 欧美日韩成人在线一区二区| 丝袜脚勾引网站| 日本-黄色视频高清免费观看| 亚洲高清免费不卡视频| 国产欧美亚洲国产| 啦啦啦中文免费视频观看日本| 人妻一区二区av| 成人18禁高潮啪啪吃奶动态图| 欧美变态另类bdsm刘玥| 国产精品免费大片| 在线观看免费视频网站a站| 2018国产大陆天天弄谢| 在线观看一区二区三区激情| 日韩不卡一区二区三区视频在线| 韩国高清视频一区二区三区| 91午夜精品亚洲一区二区三区| 最近中文字幕高清免费大全6| 亚洲av国产av综合av卡| 多毛熟女@视频| 亚洲国产看品久久| av免费在线看不卡| 十八禁高潮呻吟视频| 国产精品无大码| 久久久精品区二区三区| 免费高清在线观看视频在线观看| 精品一区二区三区视频在线| 国产成人精品久久久久久| 亚洲国产最新在线播放| 女性生殖器流出的白浆| 日本av手机在线免费观看| 蜜桃在线观看..| 亚洲精品国产色婷婷电影| 日韩一区二区视频免费看| 黑人高潮一二区| 丝袜喷水一区| av片东京热男人的天堂| 男人舔女人的私密视频| 亚洲天堂av无毛| 99热这里只有是精品在线观看| 这个男人来自地球电影免费观看 | 久久精品人人爽人人爽视色| 国产av国产精品国产| 一本一本久久a久久精品综合妖精 国产伦在线观看视频一区 | 老司机影院毛片| 少妇猛男粗大的猛烈进出视频| 国产探花极品一区二区| 国产视频首页在线观看| 一区二区日韩欧美中文字幕 | 水蜜桃什么品种好| 日本黄大片高清| 成人无遮挡网站| 亚洲在久久综合| 欧美性感艳星| 伦精品一区二区三区| 丰满迷人的少妇在线观看| 久久毛片免费看一区二区三区| av免费在线看不卡| 中文字幕免费在线视频6| 欧美日韩视频精品一区| 亚洲av综合色区一区| 亚洲熟女精品中文字幕| 婷婷成人精品国产| 香蕉国产在线看| 婷婷成人精品国产| 亚洲熟女精品中文字幕| 亚洲国产欧美日韩在线播放| 国产精品国产三级专区第一集| 免费观看无遮挡的男女| 女人久久www免费人成看片| 精品一区在线观看国产| 伊人久久国产一区二区| 日韩精品有码人妻一区| 亚洲精品456在线播放app| 波多野结衣一区麻豆| 97人妻天天添夜夜摸| 这个男人来自地球电影免费观看 | 国产精品 国内视频| 水蜜桃什么品种好| 日韩av在线免费看完整版不卡| 熟女人妻精品中文字幕| 久久免费观看电影| 亚洲精品第二区| 大香蕉久久成人网| 亚洲精品中文字幕在线视频| 欧美人与性动交α欧美精品济南到 | 久久国产精品大桥未久av| videossex国产| 国产av码专区亚洲av| 欧美国产精品va在线观看不卡| 男的添女的下面高潮视频| 51国产日韩欧美| av播播在线观看一区| 国产精品国产三级国产专区5o| 国产精品成人在线| 黄色 视频免费看| 国产熟女午夜一区二区三区| 成年av动漫网址| 久久久久久伊人网av| tube8黄色片| 大片电影免费在线观看免费| 人妻 亚洲 视频| 男女高潮啪啪啪动态图| 尾随美女入室| av国产精品久久久久影院| 亚洲,欧美,日韩| 久久精品久久精品一区二区三区| 久久鲁丝午夜福利片| 午夜福利视频精品| 插逼视频在线观看| 亚洲国产色片| 久久久亚洲精品成人影院| a级毛色黄片| 国产一区二区在线观看日韩| 99久久人妻综合| 最近最新中文字幕免费大全7| 青春草视频在线免费观看| 丰满乱子伦码专区| 高清视频免费观看一区二区| 啦啦啦视频在线资源免费观看| 久久国产精品大桥未久av| 大香蕉97超碰在线| 久久久亚洲精品成人影院| 久久精品久久久久久久性| 中文字幕人妻熟女乱码| 亚洲av在线观看美女高潮| 一本—道久久a久久精品蜜桃钙片| 亚洲三级黄色毛片| 国产精品一区二区在线不卡| 亚洲在久久综合| 丝袜美足系列| 成年美女黄网站色视频大全免费| 老司机影院成人| 汤姆久久久久久久影院中文字幕| 美女视频免费永久观看网站| 久久这里只有精品19| 日本欧美视频一区| 丰满乱子伦码专区| 极品少妇高潮喷水抽搐| 深夜精品福利| 一区二区三区四区激情视频| 久久国产精品大桥未久av| 最后的刺客免费高清国语| 精品熟女少妇av免费看| 人妻少妇偷人精品九色| 国产精品成人在线| 蜜桃国产av成人99| 人人妻人人澡人人看| 狠狠精品人妻久久久久久综合| 亚洲精品乱久久久久久| 精品福利永久在线观看| 男女国产视频网站| 亚洲久久久国产精品| 国产精品久久久久久av不卡| 亚洲欧美一区二区三区黑人 | 精品卡一卡二卡四卡免费| av.在线天堂| 一级毛片我不卡| 欧美最新免费一区二区三区| 高清欧美精品videossex| 亚洲国产精品一区二区三区在线| 99热全是精品| 日韩精品有码人妻一区| 国产又爽黄色视频| 午夜久久久在线观看| 老熟女久久久| 精品一区二区三区四区五区乱码 | 色婷婷久久久亚洲欧美| 国产精品国产三级专区第一集| 久久久久久久国产电影| xxx大片免费视频| 久久这里有精品视频免费| 亚洲色图综合在线观看| 成年美女黄网站色视频大全免费| 色婷婷久久久亚洲欧美|